Urocoptidae

Urocoptidae
Shell of Arangia sowerbyana (syntype at MNHN, Paris)
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
Order: Stylommatophora
Superfamily: Urocoptoidea
Family: Urocoptidae
Pilsbry, 1898[1] (1868)

Urocoptidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Urocoptoidea. [2]

The family Urocoptidae consists of 7 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

  • Urocoptinae Pilsbry, 1898 (1868) - synonym: Cylindrellidae Tryon, 1868
  • Apomatinae Paul, 1982
  • Brachypodellinae H. B. Baker, 1956
  • Eucalodiinae P. Fischer & Crosse, 1873
  • Holospirinae Pilsbry, 1946
  • Microceraminae Pilsbry, 1904 - synonyms: Johaniceraminae Jaume & de la Torre, 1972; Macroceraminae Jaume & de la Torre, 1972
  • Tetrentodontinae Bartsch, 1943

2008 taxonomy

Uit de Weerd (2008)[3] moved the Urocoptidae to the newly established superfamily Urocoptoidea based on molecular phylogeny research.[3]

The following cladogram based on Bayesian analysis of sequences of 28S ribosomal RNA shows the phylogenic relations within the Urocoptoidea.[3] There are up to 2008 genetically analyzed species of Urocoptidae and one species of Cerionidae marked in khaki color in the cladogram.[3]

Urocoptoidea

Macroceramus microdon

Microceramus pontificus

Torrecoptis holguinensis

Autocoptis menkeana

Simplicervix inornata

Brachypodella dominicensis gabbi

Dissotropis sp. nov.

Anisospira lebmanni

Archegocoptis crenata

Eucalodium sp.

Cerion striatellum

Coelostemma sp. nov.

Hendersoniella lux lux

Holospira fortisculpta

Epirobia polygyra

2012 taxonomy

Fred G. Thompson established the family Epirobiidae in 2012 for five genera of snails (Epirobia, Propilsbrya, Pectinistemma, Gyrocion, Prionoloplax), that were previously placed within Urocoptidae.[4] Data from molecular phylogeny research as well as morphology supports monophyly of Holospiridae Pilsbry, 1946 and the Eucalodiidae Crosse & Fischer, 1868.[4]

Genera

Genera in the family Urocoptidae include:

Brachypodellinae
  • Apoma H. Beck, 1837
  • Brachypodella Beck, 1837 - type genus of the subfamily Brachypodellinae[5]
  • Cyclauchen Pilsbry, 1930
  • Geoscala Pilsbry & Vanatta, 1898
  • Ischnostrophina Pilsbry, 1933
  • Mychostoma Albers, 1850
  • Simplicervix Pilsbry, 1903
Urocoptinae
  • Acrocoptis C. de la Torre & Bartsch, 1972
  • Allocoptis Thompson & Franz, 1976
  • Amphicosmia Pilsbry & Vanatta, 1898
  • Amphistemma C. de la Torre & Bartsch, 1972
  • Anafecoptis C. de la Torre & Bartsch, 1972
  • Angulicervix Pilsbry, 1933
  • Anoma Albers, 1850
  • Arangia Pilsbry & Vanatta, 1898
  • Archegocoptis Pilsbry, 1903[3]
  • Autocoptis Pilsbry, 1902[3]
  • Bactrocoptis Pilsbry, 1902
  • Badiofaux Pilsbry, 1941
  • Bialasmus C. de la Torre & Bartsch, 1972
  • Brachypodella L. Pfeiffer, 1840 (as Cylindrella)
  • Bostrichocentrum Strebel, 1879 : synonym of Holospira E. von Martens, 1860
  • Brevipedella Pilsbry, 1903
  • Callocoptis C. de la Torre & Bartsch, 1972
  • Callonia Crosse & P. Fischer, 1870
  • Capillacea C. de la Torre & Bartsch, 1972
  • Carcinostemma C. de la Torre & Jaume, 1972
  • Centralia de la Torre & Bartsch, 1972
  • Cochlodinella Pilsbry & Vanatta, 1898
  • Esochara Pilsbry & Vanatta, 1898
  • Fibricutis Pilsbry, 1903
  • Geminicoptis C. de la Torre & Bartsch, 1972
  • Gongylostoma Albers, 1850
  • Gyraxis Pilsbry, 1903
  • Heterocoptis C. de la Torre & Bartsch, 1972
  • Idiostemma Pilsbry & Vanatta, 1898
  • Insulaceramus Clench, 1967
  • Johaniceramus C. de la Torre & Bartsch, 1972
  • Levistemma C. de la Torre & Bartsch, 1972
  • Liocallonia Pilsbry, 1902
  • Liparotes Pilsbry, 1903
  • Maceo Pilsbry & Vanatta, 1898
  • Macroceramus Guilding, 1828
  • Microceramus Pilsbry & Vanatta, 1898 - type genus of the subfamily Microceraminae[5]
  • Nesocoptis Pilsbry, 1941
  • Nodulia C. de la Torre & Bartsch, 1972
  • Organocoptis C. de la Torre & Bartsch, 1972
  • Orienticoptis C. de la Torre & Bartsch, 1972
  • Paracallonia Pilsbry, 1903
  • Pfeiffericoptis C. de la Torre & Bartsch, 1972
  • Pineria Poey, 1854
  • Planostemma C. de la Torre & Bartsch, 1972
  • Pleurostemma C. de la Torre & Bartsch, 1972
  • Poecilocoptis Pilsbry, 1941
  • Poeycoptis C. de la Torre & Bartsch, 1972
  • Pseudopineria Aguayo, 1938
  • Pycnoptychia Pilsbry & Vanatta, 1898
  • Sagracoptis C. de la Torre & Bartsch, 1972
  • Sectilumen Pilsbry & Vanatta, 1898
  • Spiroceramus Pilsbry & Vanatta, 1898
  • Spirostemma Pilsbry & Vanatta, 1898
  • Steatocoptis Pilsbry, 1941
  • Strophina Mörch, 1852
  • Teneria C. de la Torre & Bartsch, 1972
  • Tenuistemma C. de la Torre & Bartsch, 1972
  • Tetrentodon Pilsbry, 1903 - type genus of the subfamily Tetrentodontinae[5]
  • Tomelasmus Pilsbry & Vanatta, 1898
  • Torrecoptis Bartsch, 1943[3]
  • Trilamellaxis C. de la Torre & Bartsch, 1972
  • Uncinicoptis C. de la Torre & Bartsch, 1972
  • Urocoptis Beck, 1837 - type genus of the family Urocoptidae[5]
Urocoptidae incertae sedis (temporary name)
  • Habeas Simone, 2013

References

  1. ↑ Pilsbry H. A. (1898). The Nautilus 11(9): 107.
  2. ↑ MolluscaBase eds. (2026). MolluscaBase. Urocoptidae Pilsbry, 1898 (1868). Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=932609 on 2026-07-02

    Taxonomy

    2005 taxonomy

    The family Urocoptidae was classified in the superfamily Orthalicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).<ref name="Bouchet 2005">Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.{{cite journal}}: CS1 maint: periodical has ISBN (link)
  3. 1 2 3 4 5 6 7 Uit De Weerd, D. R. (2008). "Delimitation and phylogenetics of the diverse land-snail family Urocoptidae (Gastropoda: Pulmonata) based on 28S rRNA sequence data: A reunion with Cerion". Journal of Molluscan Studies. 74 (4): 317–29. doi:10.1093/mollus/eyn023.
  4. 1 2 Thompson F. G. (2012). "The land snail genus Epirobia and allied genera in México and Central America, with the description of a new family, the Epirobiidae (Gastropoda, Pulmonata, Urocoptoidea)". Bulletin of the Florida Museum of Natural History 51: 167-215. PDF.
  5. 1 2 3 4 Cite error: The named reference Bouchet 2005 was invoked but never defined (see the help page).

Further reading

  • Bartsch, P. (1943). "Notes on Mexican Urocoptid mollusks". Journal of the Washington Academy of Sciences. 33 (2): 54–59.; 7 figs.
  • Carlos de la Torre. 1912. New Cuban Urocoptis of the U. cinerea group. The Nautilus, volume 26, number 4, pages 55-58, plate 6.
  • Pilsbry H. A. & Henderson J. B. jr. (February 1913). Two new Cuban Urocoptidae. The Nautilus, volume 26, number 10, pages 109-110.
  • (in Spanish) Miguel L. Jaume & Alfredo de la Torre. 1976. Los Urocoptidae de Cuba (Mollusca-Pulmonata). Série 4: Ciencias biologicas; no. 53. La Habana, Cuba: Centro de Información Cientifica y Tecnica, 122 pp.
  • Mesa Garcia, R; Jaume, ML (1982). "Zoogeography, ecology, and protection of the Urocoptidae family in Cuba (Gastropoda-Pulmonata). II". Revista Cubana de Medicina Tropical. 34 (2): 172–92. PMID 6763741.
  • Richardson C. L. 1991. Urocoptidae: catalogue of species. Tryonia 22: 1–245.
  • Tryon G. W. (1903) Manual of Conchology. Second series: Pulmonata. Volume 15. Urocoptidae. Continued by H. A. Pilsbry.