Tylototriton

Tylototriton
Tylototriton shanjing
Tylototriton umphangensis
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Amphibia
Order: Urodela
Family: Salamandridae
Subfamily: Pleurodelinae
Genus: Tylototriton
Andersson, 1871
Type species
Tylototriton verrucosus
Anderson, 1871

Tylototriton is a genus of newts commonly known as crocodile newts or knobby newts. About 38 known species are in this genus, with many species having been described recently.[1][2][3] They range from northeastern India and Nepal through Burma to northern Thailand, Laos, Vietnam, and southern China.[2]

Species

This genus currently contains around 40 described species as of April 2026:[4][5]

  • Tylototriton anguliceps Le, Nguyen, Nishikawa, Nguyen, Pham, Matsui, Bernardes, and Nguyen, 2015
  • Tylototriton anhuiensis Qian, Sun, Li, Guo, Pan, Kang, Wang, Jiang, Wu, and Zhang, 2017
  • Tylototriton asperrimus Unterstein, 1930
  • Tylototriton broadoridgus Shen, Jiang, and Mo, 2012
  • Tylototriton dabienicus Chen, Wang, and Tao, 2010
  • Tylototriton daloushanensis Zhou, Xiao, and Luo, 2022
  • Tylototriton gaowangjienensis  Huang, Xiang, Wu, Zhang, Zhang, Wang, Lan, Huang, Jiang, and Jiang, 2024
  • Tylototriton guilinensis Xiao et al., 2026[6]
  • Tylototriton hainanensis Fei, Ye, and Yang, 1984
  • Tylototriton himalayanus Khatiwada, Wang, Ghimire, Vasudevan, Paudel, and Jiang, 2015
  • Tylototriton kachinorum Zaw, Lay, Pawangkhanant, Gorin, and Poyarkov, 2019
  • Tylototriton kweichowensis Fang and Chang, 1932
  • Tylototriton liuyangensis Yang, Jiang, Shen, and Fei, 2014
  • Tylototriton lizhengchangi Hou, Zhang, Jiang, Li and Lu, 2012
  • Tylototriton maolanensis Li, Wei, Cheng, Zhang, and Wang, 2020
  • Tylototriton ngarsuensis Grismer, Wood, Quah, Thura, Espinoza, Grismer, Murdoch, and Lin, 2018
  • Tylototriton ngoclinhensis Phung, Pham, Nguyen, Ninh, Nguyen, Bernardes, Le, Ziegler & Nguyen, 2023
  • Tylototriton notialis Stuart, Phimmachak, Sivongxay, and Robichaud, 2010
  • Tylototriton panhai Nishikawa, Khonsue, Pomchote, and Matsui, 2013
  • Tylototriton panwaensis Grismer, Wood, Quah, Thura, Espinoza, and Murdoch, 2019
  • Tylototriton pasmansi Bernardes, Le, Nguyen, Pham, Pham, Nguyen, and Ziegler, 2020
  • Tylototriton phukhaensis Pomchote, Khonsue, Thammachoti, Hernandez, Peerachidacho, Suwannapoom, Onishi, and Nishikawa, 2020
  • Tylototriton podichthys Phimmachak, Aowphol, and Stuart, 2015
  • Tylototriton pseudoverrucosus Hou, Gu, Zhang, Zeng, and Lu, 2012
  • Tylototriton pulcherrimus Hou, Zhang, Li, and Lu, 2012
  • Tylototriton shanjing Nussbaum, Brodie, and Yang, 1995
  • Tylototriton shanorum Nishikawa, Matsui, and Rao, 2014
  • Tylototriton sini Lyu, Wang, Zeng, Zhou, Qi, Wan, and Wang, 2021
  • Tylototriton sparreboomi Bernardes, Le, Nguyen, Pham, Pham, Nguyen, and Ziegler, 2020
  • Tylototriton taliangensis Liu, 1950
  • Tylototriton thaiorum Poyarkov, Nguyen, and Arkhipov, 2021
  • Tylototriton tongziensis Li, Liu, Shi, Wei, and Wang, 2022
  • Tylototriton umphangensis Pomchote, Peerachidacho, Hernandez, Sapewisut, Khonsue, Thammachoti, and Nishikawa, 2021
  • Tylototriton uyenoi Nishikawa, Khonsue, Pomchote, and Matsui, 2013
  • Tylototriton verrucosus Anderson, 1871
  • Tylototriton vietnamensis Böhme, Schöttler, Nguyen, and Köhler, 2005
  • Tylototriton vietnamirabilis Ong AV, Phan TQ, Hoang CV, Nguyen MHT, Nguyen TT, Ziegler T, Nguyen TQ, Pham CT, 2026
  • Tylototriton wenxianensis Fei, Ye, and Yang, 1984
  • Tylototriton wufengensis Li et al., 2026[7]
  • Tylototriton yangi Hou, Zhang, Zhou, Li, and Lu, 2012
  • Tylototriton zaimeng Decemson, Lalremsanga, Elangbam, Vabeiryureilai, Shinde, Purkayastha, Arkhipov, Bragin & Poyarkov, 2023
  • Tylototriton ziegleri Nishikawa, Matsui, and Nguyen, 2013

Phylogeny

The following phylogeny of Tylototriton is from Nishikawa, Matsui & Rao (2014).[8]

Tylototriton

References

  1. ↑ Max Sparreboom (2013). "Tylototriton Anderson, 1871". Salamanders of the Old World. Retrieved 14 April 2013.
  2. 1 2 Frost, Darrel R. (2021). "Tylototriton Anderson, 1871". Amphibian Species of the World 6.1, an Online Reference. American Museum of Natural History. Retrieved 19 November 2021.
  3. ↑ Nishikawa, Kanto; Masafumi Matsui; Tao Thien Nguyen (2013). "A new species of Tylototriton from northern Vietnam (Amphibia: Urodela: Salamandridae)". Current Herpetology. 32: 34–49. doi:10.5358/hsj.32.34. S2CID 59036508.
  4. ↑ Phung, Trung My; Pham, Cuong The; Nguyen, Truong Quang; Ninh, Hoa Thi; Nguyen, Huy Quoc; Bernardes, Marta; Le, Son Thanh; Ziegler, Thomas; Nguyen, Tao Thien (2023-07-03). "Southbound – the southernmost record of Tylototriton (Amphibia, Caudata, Salamandridae) from the Central Highlands of Vietnam represents a new species". ZooKeys (1168): 193–218. Bibcode:2023ZooK.1168..193P. doi:10.3897/zookeys.1168.96091. ISSN 1313-2970. PMC 10848839. PMID 38328626.
  5. ↑ Ong, An Vinh; Phan, Tien Quang; Hoang, Chung Van; Nguyen, Mai Hong Thi; Nguyen, Tao Thien; Ziegler, Thomas; Nguyen, Truong Quang; Pham, Cuong The (2026-04-08). "A new species of Tylototriton (Amphibia, Salamandridae) from Phu Xai Lai Leng Mountain, Nghe An Province, Vietnam". ZooKeys (1276): 285–305. Bibcode:2026ZooK.1276..285O. doi:10.3897/zookeys.1276.173848. ISSN 1313-2970. PMC 13084263. PMID 42004729.
  6. ↑ Xiao; Chen; Wu; Pan; Chen; Yu; Huang (2026). "A new species of genus Tylototriton (Caudata, Salamandridae) from northeastern Guangxi, China". Zoosystematics and Evolution. 102 (2): 671–686. Bibcode:2026ZooEv.102..671X. doi:10.3897/zse.102.184792.
  7. ↑ Li, Shize; Shi, Shengchao; Liu, Jing; Luo, Zhonghao; Wang, Jiajun; Liao, Linhong; Wang, Yeqing; Gong, Renhu; Wu, Jun; Wang, Bin (2026). "Description of a new species of the Asian newt genus Tylototriton Anderson, 1871 (Urodela, Salamandridae) from central China". Zoosystematics and Evolution. 102 (1): 181–197. doi:10.3897/zse.102.173283.
  8. ↑ Nishikawa, Matsui & Rao, 2014 : A new species of Tylototriton (Amphibia: Urodela: Salamandridae) from central Myanmar. Natural History Bulletin of the Siam Society, vol. 60, no. 1, p. 9–22.