Patriomanidae

Patriomanidae; Temporal range: Da. S T Ypr. Lut. B Pr. Rup. Ch. middle to late Eocene
	Artist reconstruction of; Patriomanis americana; compared to the size of an adult human.
	illustration of Cryptomanis gobiensis; right femur bone
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Infraclass:	Placentalia
Order:	Pholidota
Suborder:	Eupholidota
Superfamily:	Manoidea
Family:	†Patriomanidae; Szalay & Schrenk, 1998
Type genus
	†Patriomanis; Emry, 1970
Genera
	†Cryptomanis; †Patriomanis;

Patriomanidae ("fathers of pangolins") is an extinct family of pangolins from superfamily Manoidea that includes two extinct genera Patriomanis and Cryptomanis.^[2]^[3] Their fossils are found in Asia and North America.

Taxonomy

Family: †Patriomanidae Szalay & Schrenk, 1998 Genus: †Cryptomanis Gaudin, Emry, and Pogue, 2006 †Cryptomanis gobiensis Gaudin, Emry, and Pogue, 2006 Genus: †Patriomanis Emry, 1970 †Patriomanis americana Emry, 1970 (american pangolin)

Phylogenetic position of family Patriomanidae within order Pholidota based on Rose (2026.) study:^[4]

outhgroup

Pholidotamorpha

Pholidota

Eupholidota

†Eomanoidea

Manoidea

Manidae

?

†Patriomanidae

sensu stricto

(Pholidota [sensu lato])

↑ Frederick S. Szalay and Friedemann Schrenk (1998) "The Middle Eocene Eurotamandua and a Darwinian phylogenetic Analysis." Kaupia 7, pp. 97-186
↑ Gaudin, Timothy (2009). "The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis" (PDF). Journal of Mammalian Evolution. 16 (4). Heidelberg, Germany: Springer Science+Business Media: 235–305. doi:10.1007/s10914-009-9119-9. Archived from the original (PDF) on 2015-09-25. Retrieved 2020-12-18.
↑ Kondrashov, Peter; Agadjanian, Alexandre K. (2012). "A nearly complete skeleton of Ernanodon (Mammalia, Palaeanodonta) from Mongolia: morphofunctional analysis". Journal of Vertebrate Paleontology. 32 (5): 983–1001. Bibcode:2012JVPal..32..983K. doi:10.1080/02724634.2012.694319. ISSN 0272-4634.
↑ Rose, K. D.; Rabenstein, R.; Habersetzer, J.; Gaudin, T. J. (2026). "Anatomy of Eurotamandua joresi (Mammalia) from Grube Messel, Germany, based on computed tomography, and implications for its relationships. Part I: Background and forelimb". Palaeobiodiversity and Palaeoenvironments. doi:10.1007/s12549-025-00686-x.