Hackelia

Hackelia
Hackelia diffusa
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Embryophytes
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Asterids
Order: Boraginales
Family: Boraginaceae
Subfamily: Boraginoideae
Genus: Hackelia
Opiz (1839)
Type species
Hackelia deflexa
Opiz
Species

About 54 (see text)

Synonyms[1]

Austrocynoglossum Popov ex R.Mill. (1989)

Hackelia (stickseeds) is a genus of plants in the borage family, Boraginaceae. It includes 54 species found in North America, western South America, temperate Eurasia, and Australia.[1] 12 species are native to California.[2]

The genus was named after Josef Hackel, a Czech botanist. The common name, stickseed, refers to the tendency of the barbed nutlets to stick to animal fur.[3]

Species

54 species are currently accepted:[1]

  • Hackelia amethystina J.T.Howell – amethyst stickseed – California
  • Hackelia andicola (K.Krause) Brand – Peru
  • Hackelia bella (J.F.Macbr.) I.M.Johnst. – greater showy stickseed – northwestern California and southwestern Oregon
  • Hackelia besseyi (Rydb.) J.L. Gentry – Bessey's stickseed – Arizona, Colorado, New Mexico and Texas
  • Hackelia bhutanica R.R.Mill – eastern Nepal to northwestern Arunachal Pradesh
  • Hackelia brachytuba (Diels) I.M.Johnst. – Nepal to central China
  • Hackelia brevicula (Jeps.) J.L.Gentry – Poison Canyon stickseed – eastern California
  • Hackelia californica (A.Gray) I.M.Johnst. – California stickseed – northern California and west-central Oregon
  • Hackelia ciliata (Douglas ex Lehm.) I.M.Johnst. – British Columbia, Idaho, and Washington
  • Hackelia cinerea (Piper) I.M.Johnst. – gray stickseed – Washington, Idaho, and Montana
  • Hackelia cronquistii J.L.Gentry – Cronquist's stickseed – Oregon and Idaho
  • Hackelia cusickii (Piper) Brand – Cusick's stickseed – Oregon, northern California, and northwestern Nevada
  • Hackelia davisii Cronquist – Idaho
  • Hackelia deflexa (Wahlenb.) Opiz – northern Eurasia and northern North America
  • Hackelia difformis (Y.S.Lian & J.Q.Wang) Riedl – south-central China and Tibet
  • Hackelia diffusa (Douglas ex Lehm.) I.M.Johnst. – Spreading stickseed – British Columbia, Washington, Oregon
  • Hackelia floribunda (Lehm.) I.M.Johnst. – manyflower stickseed – western and central Canada and United States
  • Hackelia gracilenta I.M.Johnst. – Colorado
  • Hackelia heliocarpa (Brand) Brand – Mexico (Chihuahua)
  • Hackelia hendersonii (Piper) Brand – Washington, Oregon, and Montana
  • Hackelia hintoniorum (B.L.Turner) Sutorý – Mexico (Oaxaca)
  • Hackelia hirsuta (Wooton & Standl.) I.M.Johnst. – New Mexico
  • Hackelia hispida I.M.Johnst. – showy stickseed – Washington, Oregon, Idaho
  • Hackelia ibapensis L.M.Shultz & J.S.Shultz – Utah
  • Hackelia latifolia (R.Br.) Dimon & M.A.M.Renner – eastern Australia and Tasmania[4]
  • Hackelia leonotis I.M.Johnst. – northeastern and central Mexico
  • Hackelia macrophylla (Brand) I.M.Johnst. – northern Pakistan and western Himalaya
  • Hackelia meeboldii Brand – western Himalaya
  • Hackelia mexicana (Schltdl. & Cham.) I.M.Johnst. – Mexico to Venezuela and Peru
  • Hackelia micrantha (Eastw.) J.L.Gentry – Jessica sticktight – western Canada and United States
  • Hackelia mundula (Jeps.) Ferris – pink stickseed – southwestern Oregon to central California
  • Hackelia murgabica Czukav. – Tajikistan
  • Hackelia nervosa (Kellogg) I.M.Johnst. – Sierra stickseed – California and northwestern Nevada
  • Hackelia obtusifolia R.R.Mill – Bhutan
  • Hackelia ophiobia R.L.Carr – southeastern Oregon, southwestern Idaho, and northern Nevada
  • Hackelia parviflora (K.Krause) Brand – Peru (Junín)
  • Hackelia patens (Nutt.) I.M.Johnst. – spotted stickseed – west-central United States
  • Hackelia pinetorum (Greene ex A.Gray) I.M.Johnst. – Arizona, New Mexico, Texas, and northern Mexico
  • Hackelia popovii Czukav. – Tajikistan
  • Hackelia rattanii (Brand) Brand – Mexico (Baja California)
  • Hackelia revoluta (Ruiz & Pav.) I.M.Johnst. – Colombia to northwestern Argentina
  • Hackelia sessilifructa (Y.S.Lian & J.Q.Wang) Ovczinnikova – Xinjiang
  • Hackelia setosa (Piper) I.M.Johnst. – bristly stickseed – northern California and southwestern Oregon
  • Hackelia sharsmithii I.M.Johnst. – Sharsmith's stickseed – central California and central Nevada
  • Hackelia skutchii I.M.Johnst. – Guatemala and Costa Rica
  • Hackelia stewartii I.M.Johnst. – northern Pakistan and western Himalaya
  • Hackelia stricta I.M.Johnst. – southwestern Mexico to Veracruz
  • Hackelia suaveolens (R.Br.) Dimon & M.A.M.Renner – eastern and southern Australia and Tasmania[4]
  • Hackelia taylorii Harrod, Malmquist & R.L.Carr – Washington
  • Hackelia torva (Dimon & M.A.M.Renner) Ovchinnikova – Australia
  • Hackelia uncinata (Royle ex Benth.) C.E.C.Fisch. – northeastern Pakistan to Himalaya and southern China
  • Hackelia ursina (Greene ex A.Gray) I.M.Johnst. – southeastern Arizona, southwestern New Mexico, and northern Mexico (Sonora)
  • Hackelia velutina (Piper) I.M.Johnst. – velvet stickseed – north-central and central California and northwestern Nevada
  • Hackelia venusta (Piper) H.St.John – lesser showy stickseed – central Washington
  • Hackelia virginiana (L.) I.M.Johnst. – beggar's lice – central and eastern United States and eastern Canada

References

  1. 1 2 3 Hackelia Opiz. Plants of the World Online. Retrieved 17 June 2023.
  2. ↑ Calflora Database: Hackelia species index in California. Calflora. Retrieved 17 June 2023.
  3. ↑ Taylor, Ronald J. (1994) [1992]. Sagebrush Country: A Wildflower Sanctuary (rev. ed.). Missoula, MT: Mountain Press Pub. Co. p. 24. ISBN 0-87842-280-3. OCLC 25708726.
  4. 1 2 Richard J. Dimon; Matt A. M. Renner (2017). "A new Cynoglossum species, and transfers to Hackelia in eastern Australian Boraginaceae" (PDF). Australian Systematic Botany. 30 (2): 113. doi:10.1071/sb17004. ISSN 1030-1887. Wikidata Q93466754.
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