Höganäs Formation

Höganäs Formation
Stratigraphic range: Rhaetian-Hettangian
~
Historical photo of one of the main outcrops, Norra Albert mine
TypeGeological formation
Unit ofÄngelholm Trought
Sub-unitsBjuv, Helsingborg & Vallåkra members
UnderliesRya Formation
OverliesKågeröd Formation
Thickness200 m
Lithology
PrimarySandstone
OtherShale, coal
Location
Coordinates56°06′N 12°42′E / 56.1°N 12.7°E / 56.1; 12.7
Approximate paleocoordinates44°36′N 8°30′E / 44.6°N 8.5°E / 44.6; 8.5
RegionSkåne
CountrySweden
Type section
Named byTroedsson[1]
Höganäs Formation is located in Sweden
Höganäs Formation
Höganäs Formation (Sweden)

The Höganäs Formation is a Late Triassic to Early Jurassic geologic formation in Skåne, Sweden. The formation is mostly known for its incredible flora collection from the Bjuv member, composed of over 110 species, and also includes several vertebrate remains, such as fishes, amphibians and dinosaur tracks & remains, although none have yet been referred to a specific genus.[2] It´s regional equivalent is the Zagaje Formation in Poland.[3]

Description

Coal seams of the Höganas Formation

The Höganas Formation was deposited in what is now the southern part of Sweden Skåne. The formation straddles the Triassic-Jurassic boundary and has provided evidence of widespread mire-forest deterioration that began in the latest Rhaetian.[4] The upper part of the Höganas Formation is correlated with the Rønne Formation of Bornholm, which is administered by Denmark. The Hóganas Formation is rich in coal and has provided many fossil flora.[5]

Vallåkra Member

This member represent the basal part of the Formation, that represents a transition between the terrestrial Kågeröd Formation and the delta dominated coal layers of the Bjuv Member, being made by poorly stratified clay & claystone, mostly green in coloration.[6] The sediments of this member are exposed mostly in the quarry of the town of the same name, where is subdivided on the next levels: 6 m of mixed layers, whose composition includes material seen both on the Kågeröd Formation and this member (red clay and 5 m light clay, 3 m argillaceous sandstone and l m dark clay and several layers composed by Spherosiderite).[6] The uppermost levels are covered by the coal seams of the Bjuv Member. This member has been considered as of Middle and Upper Rhaetian in age.[6][7]

Bjuv Member

Reconstruction of the estuarine ecosystem with Palaeoxyris

The second member is the Bjuv Member, known as the most fossiliferous one, that was known originally as the Gruv Member (gruva= mine).[7][8] Coal and clay have been mined in this area since the fifteenth century, being this intensive mining one of the main reasons the fossiliferous layers have provided such a diverse flora. The Bjuv member is divided on two different seams, the B seam, that overlies the older member and the A seam, that marks the limit with the Helsingborg Member.[7] This layer has a thickness of around 37.22 and 45.72 m in bore No. 256 (stored at the Höganäs Company in Bjuv) from the Bjuv area.[6] The layer is dominated by coarse arenaceous sediments, specially between Billesholm and Höganäs, parallel to the Söderåsen and Kullaberg horsts.[8] Towards the southern part of this area the deposition changes, with an increased abundance of argillaceous contents, specially at places like Skromberga.[8] The age of this member is considered to be Latest Rhaetian and maybe Lower Hettangian. Thus the member records the Triassic–Jurassic extinction, what is seen in the plant oscillations.[9] The Plant fossils found at this layer are one of the main fossil material of the whole Mesozoic in Sweden, composed by more than 110 species, one of the most complete floras of the same age from any location.[7][10] The main reason for this datation where suggested due to the presence of the plants Lepidopteris ottonis and Thaumatopteris shenki, as well palynological samples that suggested a Middle Rhaetian age (the Rhaetipollis-Limbosporites Miospore Zone).[11] Latter works have fount that the Dinosaur Footprint layers spam between the Rhaetian-Hettangian boundary.[12][13] Beyond the exceptional flora, the member is also known for fossils of Molluscs, arthropods and fish remains.[7] In the uppermost sequence the deposition changes from a coastal swamp with abundant evidence of plant colonization to fully marine conditions, evidenced by dinoflagellate cysts, recording thus the Rhaetian maximum flooding.[14]

Helsingborg Member

The Uppermost member of the formation is entirely of Hettangian age, and is the last layer before the start of the deposition of the younger dominant marine Rya Formation.[6] This member is dominated by Deltaic sediments, along several marine interealatians are frequent, with deposition of coarse, in some cases arkosic sandstones in between more fine-grained deposits may indicate Early Kimmerian tectonic movements.[15] The member has a maximunn thickness of 215 m and is divided in 2 parts: Höllviken No. 2 indicated that the lower part starts with up to 12 m of kaolinitic clay succeeded by a 15–20 m thick sequence of cross-bedded arkoses and light-coloured clays, being called Boserup beds or Thaumatopteris schenki zone (Pinuspollenites-Trachysporites Miospore Zone).[16][17] This beds are known from Boserup, but also from the central part of Helsingborg, the Köpinge No. 3 bore and even extent to the Höganäs region (northwest to Eslöv in the south).[7] This beds are overlain by some metres of arenaceous clays, iron claystone and calcareous sandstone with few intercalation of a coal seam (known as Tågaborg coal seam), where floral remains are also recovered such as Dictyophyllum nilssoni and Equisetites spp.[7][13] Then the layers turn into 5 m of argillaceous sandstone and 5 m slightly arenaceous clays, with a last 1 m of calcareous sandstone, on a sequence known from cliff series at Laröd-Sofiero at the north of Helsingborg, with plant remains such as Nilssonia polymorpha and Dictyophyllum nilssoni, referred to the Pinuspallenites-Trachysporites zone.[6] It also appears on Rosendal, Fleninge, Farhult and Klappe.[6] Helsingborg (Vossbäumer I 969). There is a coal seam, Tinkarpsflötsen, in the middle part of the sequence. The upper part of the Helsingborg Member starts with a 28 m thick sandstone, with a section known as Fleninge beds, that represent layers where marine ingressions were more common, with the last layers of the formation made up of sandstone, claystone and several minor coal seams.[6]

Fossil content

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Ichnofossils

Genus Species Location Material Made By
Aulichnites[18]
  • A. isp
  • Bjuv
  • Hyllinge
  • Farhult
  • Klappe
  • Oregården
  • Rosendal
Dwelling structures
  • Gastropods
Diplichnites[18]
  • D. ispp.
  • Bjuv
  • Hyllinge
  • Farhult
  • Klappe
  • Oregården
  • Rosendal
Hypichnial trackway
Diplocraterion[18]
  • D. parallelum
  • D. ispp.
  • Bjuv
  • Hyllinge
  • Farhult
  • Klappe
  • Oregården
  • Rosendal
U-Shaped Burrows
Lockeia[18]
  • L. siliquaria
  • Bjuv
  • Hyllinge
  • Farhult
  • Klappe
  • Oregården
  • Rosendal
Dwelling traces
  • Bivalves
Monocraterion[18]
  • M. tentaculatum
  • Bjuv
  • Hyllinge
  • Farhult
  • Klappe
  • Oregården
  • Rosendal
U-Shaped Burrows
  • Echiurans
  • Enteropneustans
  • Polychaeta
  • Sipuncula
Planolites[18]
  • P. annularis
  • P. montanus
  • Bjuv
  • Hyllinge
  • Farhult
  • Klappe
  • Oregården
  • Rosendal
Horizontal burrows
  • Annelids
  • Bivalves
  • Crustaceans
Rhizocorallium[18]
  • R. ispp.
  • Bjuv
  • Hyllinge
  • Farhult
  • Klappe
  • Oregården
  • Rosendal
Tubular Fodinichnia
  • Annelids
  • Crustaceans
  • Insects
Skolithos[18]
  • S. isp.
  • Bjuv
  • Hyllinge
  • Farhult
  • Klappe
  • Oregården
  • Rosendal
Sac/Bottle shaped burrows
  • Annelids
  • Crustaceans
  • Insects
Teichichnus[18]
  • T. isp.
  • Bjuv
  • Hyllinge
  • Farhult
  • Klappe
  • Oregården
  • Rosendal
Dwelling traces
  • Echiurans
  • Holothurians
  • Polychaetes

Annelida

Genus Species Stratigraphic position Member Material Notes Images

Burejospermum[19]

  • B. crassitestum
  • Bjuv
  • Bjuv member

Cocoons

Freshwater Clitellata Cocoons (Oligochaeta and Hirudinea)

Example of leech cocoon
Placobdella, example of leech

"Cycadeospermum"[19]

  • "C." sp.
  • Bjuv
  • Bjuv member

Cocoons

Dictyothylakos[19]

  • D. pesslerae
  • D. sp.
  • Bjuv
  • Bjuv member

Cocoons

Bivalves

Genus Species Location Stratigraphic position Material Notes Images
Avicula[20] A. nilssoni
  • Rosendal
  • Farhult
  • Oregården
  • Klappe
  • Bjuv member
  • Helsingborg member
Shells An Oyster member of Pteriidae

Cardinia[6]

C. ingelensis

  • Rosendal
  • Farhult
  • Oregården
  • Klappe
  • Bjuv member
  • Helsingborg member

Shells

A Clam member of Cardiniidae inside Carditida.

Cercomya[21] C. carlsoni
  • Bjuv
  • Bjuv member
Shells A piddock member of Ceromyidae.

Eutrapetzium[6]

E. pullastra

  • Rosendal
  • Farhult
  • Oregården
  • Klappe
  • Bjuv member
  • Helsingborg member

Shells

An Oyster member of Ostreidae. The local material assigned to this genus is now lost

Gervillia[20] G. angelini
  • Rosendal
  • Farhult
  • Oregården
  • Klappe
  • Bjuv member
  • Helsingborg member
Shells An Oyster member of Bakevelliidae

Liostrea[6][20]

L. hisingeri

  • Bjuv
  • Rosendal
  • Farhult
  • Oregården
  • Klappe
  • Helsingborg
  • Bjuv member
  • Helsingborg member

Shells

An Oyster member of Ostreidae . The most common Bivalve found locally.

Example of Liostrea specimens
Mesodesma[20] M. germari
  • Rosendal
  • Farhult
  • Oregården
  • Klappe
  • Bjuv member
  • Helsingborg member
Shells A Clam member of Mesodesmatidae inside Venerida.

Modiolus[6]

M. hoffinanni

  • Rosendal
  • Farhult
  • Oregården
  • Klappe
  • Helsingborg
  • Bjuv member
  • Helsingborg member

Shells

A Mussel member of Mytilidae.

Example of extant Modiolus
Mya[20]
  • M. ovalis
  • M. elongatus
  • Rosendal
  • Farhult
  • Oregården
  • Klappe
  • Bjuv member
  • Helsingborg member
Shells A Clam member of Myidae inside Myida.
Pleuromya[20] P. aquarum
  • Rosendal
  • Farhult
  • Oregården
  • Klappe
  • Bjuv member
  • Helsingborg member
Shells A Clam member of Pleuromyidae inside Myida.
Protocardia[20]
  • P. eicaldi
  • P. praecursor
  • P. suecica
  • Rosendal
  • Farhult
  • Oregården
  • Klappe
  • Bjuv member
  • Helsingborg member
Shells A Clam member of Protocardiinae inside Carditida.
Venerupis[20]
  • V. elongala
  • V. herberti
  • Rosendal
  • Farhult
  • Oregården
  • Klappe
  • Bjuv member
  • Helsingborg member
Shells A Clam member of Veneridae inside Venerida.
Example of extant Venerupis

Insecta

Genus Species Location Stratigraphic position Material Notes Images

Angelinella[22]

A. angelini

  • Höganäs

Bjuv member

Elytrons

A beetle, incertae sedis Archostemata

Heeriaopsis[22]

H. laevigatum

  • Höganäs

Bjuv member

Elytrons

A beetle, incertae sedis Archostemata

Nannocurculionites[22]

N. carlsoni

  • Bjuv

Bjuv member

Elytrons

A beetle, incertae sedis Archostemata. Originally identified as Curculionites carlsoni

Parabuprestites[22]

P. rugulosus

  • Bjuv

Bjuv member

Elytrons

A Beetle, member of the family Cupedidae. Originally identified as Buprestites rugulosus

Pseudocarabites[22]

P. deplanatus

  • Bjuv

Bjuv member

Elytrons

A Beetle, member of the family Carabidae. Originally identified as Carabites deplanatus.

Pseudohydrophilites[22]

P. nathorsti

  • Bjuv

Bjuv member

Elytrons

A Beetle, member of the family Schizocoleidae. Originally identified as Hydrophilites nathorsti.

Fish

Genus Species Location Stratigraphic position Material Notes Images
Hybodus[23] H. sp.
  • Helsingborg
Helsingborg member
  • Isolated Teeth
A freshwater/marine member of Hybodontidae.
Hybodus reconstruction
Hyllingea[24] H. swanbergi
  • Hyllinge
Bjuv member
  • Incomplete Specimens
  • Isolated Scales
A freshwater member of the family Acrolepidae.
Acrolepis, a relative of Hyllingea
Lissodus[23] L. sp.
  • Helsingborg
Helsingborg member
  • Isolated Teeth
A freshwater/marine member of Lonchidiidae.
Palaeoxyris[25] P. muensteri
  • Bjuv
  • Helsingborg
  • Bjuv member
  • Helsingborg member
  • Shark eggs Capsules
A freshwater shark egg capsule, likely produced by members of Hybodontidae. The eggs are found in close association with Neocalamites stems (An ideal substrate to attach them).[25]
Parvodus[23] P. sp.
  • Helsingborg
Helsingborg member
  • Isolated Teeth
A freshwater/marine member of Lonchidiidae.
Saurichthys[21] S. acuminatus
  • Bjuv
Bjuv member
  • Incomplete Specimens
  • Isolated Scales
A freshwater member of the family Saurichthyidae.
Saurichthys reconstruction

Scanilepis[26][27]

S. dubia

  • Bjuv
Bjuv member (Rhaetian)
  • Single Incomplete Specimen
  • Isolated Teeth
  • Isolated Scales

A freshwater member of the family Scanilepididae.

Semionotus[21] S. nilssoni
  • Bjuv
Bjuv member
  • Incomplete Specimens
  • Isolated Scales
A freshwater member of the family Semionotidae.
Semionotus reconstruction

Amphibia

Genus Species Location Stratigraphic position Material Notes Images

Gerrothorax[28]

G. pulcherrimus

  • coal-mine at Bjuv
Bjuv member
  • LU 3176T, LU 3176aT, LU 3176bT (holotype), postcrania including interclavicles and clavicles, vertebrae and osteoderms
  • NR B2–4, partial skull
  • NR B18a, b, c, d, e, postcranial remains

An amphibian, member of the family Plagiosauridae. Represents the only major tetrapod macrofossil from this layers.

Gerrothorax reconstruction

Dinosauria

Tracks discovered in the Höganäs Formation have been assigned to the ichnogenus Grallator (Eubrontes) cf. giganteus, which were discovered in Rhaetian strata, and Grallator (Eubrontes) soltykovensis, which were discovered in Hettangian strata.[29] A few of the tracks were taken to museums, but most of them disappeared in natural floodings.[29]

After the end of a seven-year excavation project at Billesholm, researchers announced on 13 October 2022 the identification of seven dinosaur species based on fossil remains, out of which three species were new to science.[30] The fossils are yet to be analyzed and scientifically published.[31]

Genus Species Location Stratigraphic position Material Notes Images

Eubrontes[29][32]

  • E. giganteus
  • Gustaf Adolf mine
  • Helsingborg tunnel
  • Höganäs
  • Bjuv
  • Bjuv Member (Rhaetian)

Footprints

Theropod tracks, type member of the ichnofamily Eubrontidae, incertae sedis inside Theropoda. Eubrontes tracks were likely made by Dilophosaurus-like theropods.

Cast of Eubrontes footprint

Kayentapus[29][33][12]

  • K. soltykovensis
  • Southern quarry at Vallåkra
  • Helsingborg Member (Hettangian)

Footprints

Theropod tracks, member of the ichnofamily Eubrontidae, incertae sedis inside Neotheropoda. The footprints are small-sized, tridactyl and relatively narrow. Likely made by Coelophysidae-like dinosaurs.

Footprint at Höganäs (Højenæs) Museum.

Theropoda? indet.[34]

Indeterminate[2]

  • Höganäs mine
  • Billesholm mine

Helsingborg member

  • 4 dorsal vertebrae
  • Vertebral column

Incertae sedis Dinosaurian remains, suggested to be from an indeterminate Theropod

Thyreophora? indet.[35]

Indeterminate

  • Gustaf Adolf mine
  • Bjuv Member (Rhaetian)

Footprints

The tracks described resemble basal Thyreorphora feet. Some of the tracks were lost.
Probable thyreophoran track from the Gustav Adolf Mine

Flora

Genus Species Stratigraphic position Member Material Notes Images
Allicospermum[36][37]
  • A. spp.
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
Seeds Ginkgoaceae Ginkgoales.
Anomozamites[36][38][39]
  • A. minor
  • A. gracilis
  • A. marginatus
  • A. angustifolius
  • A. intermedium
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Leaflets Williamsoniaceae Bennettitales.
Antherangiopsis[40]
  • A. rediviva
  • Coal-mine at Bjuv
  • Bjuv Member
Leaflets Cycadales Cycadopsida.
Antevsia[38][41]
  • A. zeilleri
  • Coal-mine at Bjuv
  • Hyllinge
  • Bjuv Member
Reproductive structure Peltaspermaceae Pteridospermatophyta.
Baiera[36]
  • B. taeniata
  • Coal-mine at Bjuv
  • Bjuv Member
  • Helsingborg Member
Isolated Leaves Ginkgoaceae Ginkgoales.
Baiera
Beania[40]
  • B. carruthersi
  • Coal-mine at Bjuv
  • Bjuv Member
Reproductive structure Cycadales Cycadopsida.
Brachyphyllum[37]
  • B. sp. cf. B. crucis
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Leave Compressions Hirmeriellaceae Pinales
Camptopteris[38][42]
  • C. spiralis
  • C. incisa
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
Isolated pinnae Matoniaceae Gleicheniales
Camptophyllum[36]
  • C. schimperi
  • Coal-mine at Bjuv
  • Bjuv Member
  • Helsingborg Member
Stems Likely Lycopodiopsida
Ctenis[38]
  • C. nilssonii
  • C. minuta
  • C. latepinnata
  • C. laxa
  • Coal-mine at Bjuv
  • Bjuv Member
  • Helsingborg Member
Leaflets Cycadales Cycadopsida.
Ctenopteris[36]
  • C. cycadea
  • Coal-mine at Bjuv
  • Bjuv Member
Isolated pinnae Polypodiaceae Polypodiales.
Cladophlebis[36][37][38]
  • C. svedbergii
  • C. scoresbyensis
  • C. cf. nebbensis
  • C. cf. spectabilis
  • C. arguta
  • C. sublobata
  • C. sewardii
  • C. divaricata
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Hyllinge
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Osmundaceae Osmundales.
Cladophlebis
Clathropteris[38]
  • C. meniscioides
  • Coal-mine at Bjuv
  • Hyllinge
  • Bjuv Member
Isolated pinnae Dipteridaceae Polypodiales.
Cycadites[36]
  • C. longiolius
  • Coal-mine at Bjuv
  • Bjuv Member
  • Helsingborg Member
Leaflets Cycadales Cycadopsida.
Cycadinocarpus[36][40]
  • C. erdmanni
  • Coal-mine at Bjuv
  • Bjuv Member
  • Helsingborg Member
Seeds Cycadales Cycadopsida.
Cycadolepis[38]
  • C. giganteus
  • C. blomquisti
  • Coal-mine at Bjuv
  • Bjuv Member
Leaflets Cycadales Cycadopsida.
Cycadocephalus[40]
  • C. sewardi
  • Coal-mine at Bjuv
  • Bjuv Member
Seeds Cycadales Cycadopsida.
Cycadospadix[40]
  • C. integer
  • Coal-mine at Bjuv
  • Bjuv Member
Seeds Cycadales Cycadopsida.
Cyparissidium[38][42]
  • C. nilssonianum
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
Fragmentary axis compressions with preserved leaves Either Cupressales or Palissyales
Czekanowskia[37]
  • C. setacea
  • C. sp.
  • Coal-mine at Bjuv
  • Bjuv Member
  • Helsingborg Member
Reproductive Organs Czekanowskiales Ginkgoales.
Danaeopsis[38]
  • D. fecunda
  • Coal-mine at Bjuv
  • Hyllinge
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Marattiidae Marattiales
Desmiophyllum[38][42]
  • D. cyclostomum
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
Isolated Leaves Incertade Sedis, maybe Palissyales
Dictyophyllum[36][38]
  • D. nilssoni
  • D. müensteri
  • D. exile
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
Isolated pinnae Dipteridaceae Polypodiales.
Dioonites[40]
  • D. spectabilis
  • Coal-mine at Bjuv
  • Bjuv Member
Leaflets Cycadales Cycadopsida.
Doratophyllum[38]
  • D. astartensis
  • D. scanicum
  • Coal-mine at Bjuv
  • Bjuv Member
Leaflets Cycadales Cycadopsida.
Elatocladus[38][42]
  • E. cephalotaxoides
  • E. sp. cf. Palissya sphenolepis
  • E. conferta
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Fragmentary axis compressions with preserved leaves Either Cupressales or Palissyales
Equisetites[36][38][42]
  • E. laevis[43]
  • E. praelongus[43]
  • E. müensteri[43]
  • E. subulatus[43]
  • E. beani[43]
  • E. scanicus[43]
  • E. mobergii[43]
  • E (Equisetostachys) nathorsti[43]
  • E (Equisetostachys) suecicus[43]
  • E. gracilis
  • E. spp
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Stems Equisetaceae Equisetales.
Ginkgoites[37][38][42]
  • G. troedsonii
  • G. obovatus
  • G. marginatus
  • G. sp. cf. G. marginatus
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Leave Compressions Ginkgoaceae Ginkgoales.
Ginkgoites
Hausmannia[38]
  • H. forchhammeri
  • H. dentata
  • H. lasciniata
  • Coal-mine at Bjuv
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Dipteridaceae Polypodiales.
Hydropterangium[38][44]
  • H. marsilioides
  • H. hyllingensis
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Hyllinge
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Caytoniaceae Pteridospermatophyta.
Komlopteris[36]
  • K. nordenskioeldii
  • Coal-mine at Bjuv
  • Bjuv Member
Isolated pinnae Corystospermaceae Pteridospermatophyta.
Lepidopteris[38][41]
  • L. ottonis
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Peltaspermaceae Pteridospermatophyta.
Lepidopteris
Lycopodites[38]
  • L. scanicus
  • Coal-mine at Bjuv
  • Hyllinge
  • Bjuv Member
Stems Lycopodiaceae Lycopodiales
Lycostrobus[45]
  • L. scottii
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Hyllinge
  • Bjuv Member
  • Helsingborg Member
Stems and cones The youngest Pleuromiales
Lycostrobus
Marattiopsis[38]
  • M. crenulatus
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Hyllinge
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Marattiidae Marattiales
Neocalamites[37][38][42]
  • N. hoerensis[43]
  • N. lehmannianus
  • N. sp.
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Stems Neocalamiteceae Equisetales
Nilssonia[36][38][46]
  • N. münsteri
  • N. acuminata
  • N. polymorpha
  • N. pterophylloides
  • N. brevis
  • N. fallax
  • N. pumila
  • N. spp.
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Hyllinge
  • Bjuv Member
  • Helsingborg Member
Leaflets Cycadeoidaceae Bennettitales.
Nilssoniopteris[38]
  • N. vittata
  • N. spp.
  • Coal-mine at Bjuv
  • Hyllinge
  • Bjuv Member
Leaflets Cycadeoidaceae Bennettitales.
Palissya[36][47]
  • P. braunii
  • P. sphenolepis
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
Cones Palissyales Pinales
Peltaspermum[38][41]
  • P. rotula
  • Coal-mine at Bjuv
  • Hyllinge
  • Bjuv Member
Reproductive structure Corystospermaceae Pteridospermatophyta.
Phlebopteris[36][38][42]
  • P. muensteri
  • P. angustiloba
  • P. polypodioides
  • P. spp.
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Matoniaceae Gleicheniales
Phlebopteris
Podozamites[36][38]
  • P. lanceolatus
  • P. distans
  • P. ovalis
  • P. angustifolius
  • P. cuspiformis
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Bjuv Member
  • Helsingborg Member
Isolated Leaves Krassiloviaceae Voltziales.
Podozamites
Ptilozamites[38][42]
  • P. blasii
  • P. carlsonii
  • P. heeri
  • P. nilssonii
  • P. sp. cf. P. nilssonii
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Hyllinge
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Corystospermaceae Pteridospermatophyta.
Pseudoctenis[38]
  • P. florinii
  • Coal-mine at Bjuv
  • Bjuv Member
Leaflets Cycadeoidaceae Bennettitales.
Pseudotorellia[37]
  • P. nordenskioeldii
  • P. spp.
  • Coal-mine at Bjuv
  • Hyllinge
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Leave Compressions Ginkgoaceae Ginkgoales.
Pterophyllum[38][39][42]
  • P. majus
  • P. kochii
  • P. aequale
  • P. subaequale
  • P. ptilum
  • P. compressum
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Hyllinge
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Leaflets Williamsoniaceae Bennettitales.
Pterygopteris[38]
  • P. angelini
  • Coal-mine at Bjuv
  • Hyllinge
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Osmundaceae Osmundales.
Rhaphidopteris[38]
  • R. (Stenopteris) spp.
  • Coal-mine at Bjuv
  • Bjuv Member
Isolated pinnae Corystospermaceae Pteridospermatophyta.
Rhizomopteris[36][38]
  • R. cruciata
  • R. schenki
  • Coal-mine at Bjuv
  • Bjuv Member
Isolated pinnae Dipteridaceae Polypodiales.
Sagenopteris[36][38][44]
  • S. nilssoniana
  • S. rhoifolia
  • S. undulata
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Hyllinge
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Caytoniaceae Pteridospermatophyta.
Sagenopteris
Schizolepis[38]
  • S. hörensis
  • Coal-mine at Bjuv
  • Bjuv Member
Ovulate Strobili Schizolepisaceae Pinaceae.
Selaginellites[38]
  • S. hallei
  • Coal-mine at Bjuv
  • Bjuv Member
Stems Selaginellaceae Selaginellales
Simplicioxylon[48]
  • S. hungaricum
  • Skromberga
  • Bjuv Member
Wood Hirmeriellaceae Pinales
Sorosaccus[37]
  • S. gracilis
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
Leave Compressions Ginkgoaceae Ginkgoales.
Sphenobaiera[37][38][42]
  • S. paucipartita
  • S. sp.
  • Coal-mine at Bjuv
  • Rögla area
  • Bjuv Member
Leave Compressions Ginkgoaceae Ginkgoales.
Stachyotaxus[38][42]
  • S. septentrionalis
  • S. elegans
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Isolated Leaves Palissyaceae Palissyales.
Stenorachis[38][40]
  • S. scanicus
  • S. solmsi
  • S. dubius
  • Coal-mine at Bjuv
  • Bjuv Member
Reproductive Organs Cycadeoidaceae Bennettitales.
Swedenborgia[36]
  • S. cryptomeroides
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Bjuv Member
  • Helsingborg Member
Isolated Leaves Krassiloviaceae Voltziales.
Thaumatopteris[36][38]
  • T. schenkii
  • T. brauniana
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Dipteridaceae Polypodiales.
Todites[38][42]
  • T. goeppertianus
  • T. sp. cf. T. williamsonii
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Rögla area
  • Bjuv Member
  • Helsingborg Member
Isolated pinnae Osmundaceae Osmundales.
Wielandiella[38][49]
  • W. punctata
  • W. angustifolia
  • Coal-mine at Bjuv
  • Helsingborg Harbour
  • Bjuv Member
  • Helsingborg Member
Female Bennetite "flower" Williamsoniaceae Bennettitales.
Wielandiella

See also

References

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  49. ↑ Pott, C. (2014). "A revision of Wielandiella angustifolia, a shrub-sized bennettite from the Rhaetian-Hettangian of Scania, Sweden, and Jameson Land, Greenland". International Journal of Plant Sciences. 175 (4): 467–499. Bibcode:2014IJPlS.175..467P. doi:10.1086/675577. S2CID 83990083. Retrieved 24 January 2022.

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