Endemic fauna of Uzbekistan

This is a list of the endemic fauna of Uzbekistan. It includes animal species currently reported only from Uzbekistan in major taxonomic databases and scientific publications. Some near-endemic species, whose ranges lie almost entirely within Uzbekistan, are also included. The list may change as taxonomic revisions are made and new distribution records become available.

Endemic animals of Uzbekistan
From the top clockwise: gecko Alsophylax emilia, tree trunk spider Hersiliola esyunini, stone loach Triplophysa ferganaensis, butterfly Melitaea acraeina, spider Palpimanus rakhimovi, gecko Teratoscincus rustamowi, wolf spider Zyuzicosa andreii, scorpion Mesobuthus zonsteini, gecko Alsophylax ferganensis, ant Alloformica flavicornis.

Context

Regional endemism

Uzbekistan is located in Central Asia, a region that includes several areas of high biodiversity. Parts of Uzbekistan fall within the Mountains of Central Asia biodiversity hotspot, as defined by the Critical Ecosystem Partnership Fund.[1] According to the World Wildlife Fund ecoregion classification, Uzbekistan has no ecoregions found exclusively within its borders and shares all six of its ecoregions with neighbouring countries. These include the Alai–Western Tian Shan steppe,[2] Gissaro–Alai open woodlands,[3] Badghyz and Karabil semi-desert,[4] Central Asian northern desert,[5] Central Asian southern desert,[6] and Central Asian riparian woodlands.[7][note 1] As a result, much of Uzbekistan's biodiversity is shared across national borders with neighbouring Central Asian countries, including Kazakhstan, Kyrgyzstan, Tajikistan, and Turkmenistan. Many species occurring in Uzbekistan are also expected to occur elsewhere in the region.

Central Asia has a high rate of endemism. For example, among different insect orders, the rate of endemism ranges from 10% to 50–60% of all recognised species.[8] Among millipedes, 90% of species are endemic to Central Asia.[9] At the same time, because biodiversity is shared among several countries, the number of species restricted to the territory of any single Central Asian state, including Uzbekistan, is comparatively limited.

Endemism in Uzbekistan

There is no officially recognised checklist of the endemic fauna of Uzbekistan. However, an unofficial preliminary checklist was compiled[10] by conservationist and Minnesota Zoo staff member Fred Swengel[11] as part of his global project Living National Treasures, which compiles data from scientific taxonomic databases (HBW/BirdLife Taxonomic Checklist, AviList, The Reptile Database, AmphibiaWeb, Catalog of Fishes, Lepidoptera Mundi, Catalogue of Life, and others) to identify endemic species for each country in the world.[12]

The region with the highest number of endemic species records in Uzbekistan is the Fergana Valley.[13] It is also the most densely populated region of the country and has been heavily transformed by agriculture.[14][15] Many endemic species were described from this area in the 1930s and 1940s and have not been observed since. Some, such as the moth Zygaena ferganae,[16][17] are considered extinct. At the same time, new species continue to be discovered in the valley, including vertebrates. In 2023 two new species of lizards were described: the Fergana even-fingered gecko (Alsophylax ferganensis) and Emilia's even-fingered gecko (Alsophylax emilia). Between 2021 and 2024, three new species of fish were also described from the valley: the Fergana stone loach (Triplophysa ferganaensis), the Sokh stone loach (Triplophysa daryoae), and the Uzbekistan stone loach (Dzihunia pseudoamudarjensis).

Large-scale environmental transformation during the Soviet period also affected species distributions within Uzbekistan. Beginning in the 1930s, extensive areas of arid and semi-arid land were irrigated and converted to agriculture, especially for cotton cultivation, through projects such as the Great Fergana Canal and the Virgin Lands campaign. These changes caused the loss and fragmentation of natural habitats.[13] Some species declined or disappeared, while others persisted in modified environments. One example is the butterfly Melitaea acraeina, described in 1886 from the Fergana Valley. It was not recorded again for many decades and was considered possibly extinct until its rediscovery in 1989 by Alexander Kreuzberg. Subsequent observations showed that the species survives in secondary habitats within agricultural landscapes, such as drainage canal slopes.[18] For some species, inclusion in this list may reflect that their remaining known populations are currently confined to Uzbekistan.

The biodiversity of Central Asia, including Uzbekistan, remains poorly studied. Faunistic and taxonomic research on the invertebrates of Uzbekistan has been especially limited. Much scientific effort historically focused on applied topics such as crop pest management and economically important species.[8][19] Many invertebrate species are known only from their original descriptions. Technically, such species may be treated as endemic in the literature, but this status may just reflect a lack of research and sampling rather than true geographic restriction. Inclusion in this list indicates that a species has so far been reported only from Uzbekistan in available sources; it does not necessarily imply strict endemism. Future research may document these species in neighbouring regions with similar environmental and climatic conditions.

Only species recognised by the Catalogue of Life are included in this list. Only taxa recognised at species rank are included; endemic subspecies are excluded.

There are no known endemic species of mammals, birds, or amphibians in Uzbekistan.[10][20] As of May 2026, the list includes 4 reptiles, 4 fish, 30 insects (including 3 butterflies and moths, 12 beetles, 4 ants, 5 bees, 2 wasps, 2 flies, 1 grasshopper, 1 webspinner), 27 arachnids (including 21 spiders, 4 scorpions, 2 harvestmen), 2 myriapods, 1 crustacean, 2 mollusks, and 5 earthworms. The total is 75 species. The list is not complete.

List

Endemic fauna of Uzbekistan
# Common name Picture Scientific name Distribution range / Note Red Data Book of Uzbekistan status IUCN Red List status (clickable) Catalogue of Life species profile

(clickable)

 Wikispecies profile

(clickable)

 iNaturalist species profile

(clickable)

Reptilia (reptiles)
1Fergana even-fingered geckoAlsophylax ferganensis IUCN: 100% range within Uzbekistan. Fergana Valley[21][22]-Critically EndangeredCoLWikispeciesiNaturalist
2Emilia's even-fingered geckoAlsophylax emiliaFergana Valley[23][24]--CoLWikispeciesiNaturalist
3Rustamov's plate-tailed geckoTeratoscincus rustamowiNear-endemic. IUCN range: Tajikistan and Uzbekistan, with over 90% of its range within Uzbekistan.[25][26] Fergana Valley.1 (EN) EndangeredEndangeredCoLWikispeciesiNaturalist
4Reticulated toad-headed agamaPhrynocephalus reticulatusNear-endemic. IUCN range: Turkmenistan and Uzbekistan, with over 90% of its range within Uzbekistan.[27] Kyzylkum Desert.-EndangeredCoLWikispeciesiNaturalist
Actinopterygii (ray-finned fishes)
5Fergana stone loachTriplophysa ferganaensisNear-endemic.Tajikistan and Uzbekistan, with over 90% of its range within Uzbekistan. Fergana Valley.[28]--CoLWikispeciesiNaturalist
6Sokh stone loachTriplophysa daryoaeFergana Valley[29][30]--CoL-iNaturalist
7Uzbekistan stone loachDzihunia pseudoamudarjensisFergana Valley[31][32]--CoL-iNaturalist
8Stone loach Dzihunia ilanDzihunia ilanNear-endemic. IUCN range: Tajikistan and Uzbekistan, with over 90% of its range within Uzbekistan. Zeravshan River.[33]-Least ConcernCoL-iNaturalist
Insecta (insects)
Orthoptera (grasshoppers, locusts, and crickets)
9Grasshopper Conophyma turkestanicumConophyma turkestanicumTurkestan Mountain Range[34][35]--CoL-iNaturalist
Embioptera (webspinners)
10Webspinner Uranembia rivkusiUranembia rivkusiKyzylkum Desert,[36][37] Navoiy Region, Qashqadaryo Region--CoLWikispeciesiNaturalist
Coleoptera (beetles)
11Longhorn beetle Neoplocaederus danilevskyiNeoplocaederus danilevskyiSurxondaryo Region[38]--CoLWikispeciesiNaturalist
12Click beetle Reitterelater kovalenkoiReitterelater kovalenkoiNuratau Mountains[39]--CoL--
13Fergana earth-boring dung beetle Lethrus bispinusLethrus bispinusFergana Valley[40][41]1 (CR) Critically Endangered-CoLWikispecies-
14Darkling beetle Colposcelis khudayberganiColposcelis khudayberganiKarakalpakstan[42]--CoL--
15Darkling beetle Colposcelis ustyurtensisColposcelis ustyurtensisKarakalpakstan[43]--CoL--
16Jewel beetle Acmaeoderella zeravshanicaAcmaeoderella zeravshanicaZeravshan Mountain Range[44]--CoL-iNaturalist
17Jewel beetle Acmaeoderella nigrentisAcmaeoderella nigrentisSamarkand Region[44]--CoL--
18Jewel beetle Sphenoptera variaSphenoptera variaUzbekistan[44]--CoL--
19Jewel beetle Sphenoptera caesiaSphenoptera caesiaUzbekistan[44]--CoL--
20Jewel beetle Sphenoptera korshinskiiSphenoptera korshinskiiUzbekistan[44]--CoL--
21Rove beetle Lesteva somsaLesteva somsaChatkal Mountain Range[45]--CoL--
22Weevil Gymnetron propinquumGymnetron propinquumKughitangtau Mountain Range[46]--CoL--
Lepidoptera (butterflies and moths)
23Nymphalid butterfly Melitaea acraeinaMelitaea acraeina Fergana Valley[47]1 (CR) Critically Endangered-CoL-iNaturalist
24Fergana smoky moth Zygaena ferganaeZygaena ferganaeFergana Valley[48]0 (EX)

Extinct

 -CoLWikispeciesiNaturalist
25Cossid moth Cossulus nikiforoviorumCossulus nikiforoviorumHisar Mountain Range[49]--CoLWikispeciesiNaturalist
Hymenoptera (wasps, bees, ants, and sawflies)
26Ant Leptanilla alexandriLeptanilla alexandriQashqadaryo Region[50][51]--CoL-iNaturalist
27Ant Alloformica flavicornisAlloformica flavicornisMountains of Tashkent Region. probably Kazakhstan[52]--CoLWikispecies-
28Ant Stenamma sogdianumStenamma sogdianumZeravshan Mountain Range[52][53]--CoL-iNaturalist
29Ant Proformica similisProformica similisQashqadaryo Region[52][54]--CoL-iNaturalist
30Bee Colletes uzbekusColletes uzbekusQurama Mountains[55]--CoL--
31Bee Andrena aheneaAndrena aheneaSamarkand Region[56]--CoLWikispeciesiNaturalist
32Bee Andrena lucidicollisAndrena lucidicollisTashkent region[57]--CoLWikispeciesiNaturalist
33Bee Andrena sordidellaAndrena sordidellaTashkent region[58]--CoLWikispeciesiNaturalist
34Bee Andrena subaenescensAndrena subaenescensSamarkand Region[59]--CoLWikispeciesiNaturalist
35Wasp Dinetus rakhimoviDinetus rakhimoviQashqadaryo Region[60]--CoL--
36Wasp Eumenes selisiEumenes selisiUzbekistan[61]--CoL--
Diptera (flies and mosquitoes)
37Hover fly Eumerus ryzhikEumerus ryzhikSurxondaryo Region[62]--CoL--
38Hover fly Eumerus ferulaeEumerus ferulaeKyzylkum Desert[63]1 (EN) Endangered-CoL--
Arachnida (arachnids)
Araneae (spiders)
39Palp-footed spider Palpimanus rakhimoviPalpimanus rakhimoviSurxondaryo Region[64]--CoLWikispeciesiNaturalist
40Palp-footed spider Palpimanus wagneriPalpimanus wagneriUzbekistan[65]--CoL-iNaturalist
41Tree trunk spider Hersiliola esyuniniHersiliola esyuniniFergana Valley[66]--CoLWikispeciesiNaturalist
42Crevice weaver spider Zaitunia minutaZaitunia minutaQurama Mountains,[67] Fergana Valley--CoL-iNaturalist
43Jumping spider Pseudomogrus tamdybulakPseudomogrus tamdybulakKyzylkum Desert, Navoi Region[68]--CoL-iNaturalist
44Jumping spider Pseudomogrus nuratausPseudomogrus nuratausNuratau Mountains[69][70]--CoL-iNaturalist
45Wolf spider Zyuzicosa andreiiZyuzicosa andreiiSurxondaryo Region[71][72]--CoLWikispeciesiNaturalist
46Wolf spider Zyuzicosa giganteaZyuzicosa giganteaSurxondaryo Region[73]--CoLWikispeciesiNaturalist
47Wolf spider Zyuzicosa baisunica Zyuzicosa baisunica Surxondaryo Region[73]--CoLWikispeciesiNaturalist
48Wolf spider Zyuzicosa fulviventrisZyuzicosa fulviventrisZeravshan Mountain Range[73]--CoLWikispeciesiNaturalist
49Wolf spider Zyuzicosa uzbekistanicaZyuzicosa uzbekistanicaSurxondaryo Region[73]--CoLWikispeciesiNaturalist
50Wolf spider Karakumosa gromoviKarakumosa gromoviSurxondaryo Region[74]--CoLWikispeciesiNaturalist
51Wolf spider Arctosa pardosinaArctosa pardosinaUzbekistan[75]--CoL-iNaturalist
52Tangle-web spider Theridion reinhardtiTheridion reinhardtiUzbekistan[76]--CoL-iNaturalist
53Tangle-web spider Theridion turanicumTheridion turanicumUzbekistan[77]--CoL-iNaturalist
54Spider Cedicoides simoniCedicoides simoniUzbekistan[78]--CoL-iNaturalist
55Crab spider Misumenops turanicusMisumenops turanicusUzbekistan[79]--CoL-iNaturalist
56Sac spider Clubiona maracandicaClubiona maracandicaUzbekistan[80]--CoL-iNaturalist
57Spider Liocranum freibergiLiocranum freibergiUzbekistan[81]--CoL-iNaturalist
58Ground spider Pterotricha argentosaPterotricha argentosaUzbekistan[82]--CoL-iNaturalist
59Funnel weaver spider Gorbiscape zarafshanicusGorbiscape zarafshanicusZeravshan Mountain Range[83]--CoLWikispeciesiNaturalist
Scorpiones (scorpions)
60Scorpion Orthochirus fetiOrthochirus fetiSurxondaryo Region[84]--CoLWikispeciesiNaturalist
61Scorpion Olivierus neniliniOlivierus neniliniQurama Mountains, Fergana Valley, probably Kyrgyzstan[85]--CoL-iNaturalist
62Scorpion Mesobuthus barszczevskiiMesobuthus barszczevskiiSurxondaryo Region[86]--CoLWikispeciesiNaturalist
63Scorpion Mesobuthus zonsteiniMesobuthus zonsteiniNavoiy Region[86]--CoL-iNaturalist
Opiliones (harvestmen)
64Harvestman Phalangium gromoviPhalangium gromoviSurxondaryo Region[87]--CoL-iNaturalist
65Harvestman Phalangium kitabensePhalangium kitabenseZeravshan Mountain Range[87]--CoL-iNaturalist
Myriapoda (millipedes and centipedes)
66Millipede Jaxartes zachvatkiniJaxartes zachvatkiniTashkent[88]--CoL--
67Centipede Taschkentia parthorumTaschkentia parthorumTashkent Region[89]--CoL--
Crustacea
68Woodlouse Turanoniscus anacanthotermitisTuranoniscus anacanthotermitisThe only genus and species in the family Turanoniscidae. Known only from Uzbekistan; inhabits termite nests [90][91]--CoLWikispeciesiNaturalist
Mollusca (mollusks)
69Valvatamnicola archangelskiiValvatamnicola archangelskiiUzbekistan[92][93]1 (CR)

Critically Endangered

 Data DeficientCoL-iNaturalist
70Kolhymorbis dildoraeKolhymorbis dildorae Uzbekistan[94]-Data DeficientCoL-iNaturalist
Annelida (segmented worms)
71Earthworm Allolobophora taschkentensisAllolobophora taschkentensisKhumsan, Karjantau mountains, Tashkent Region[95][96]--CoL--
72Greenheaded worm

Allolobophora chlorocephala

 Allolobophora chlorocephalaKhumsan, Karjantau mountains, Tashkent Region[95][97]2 (VU:R)

Vulnerable, naturally rare

 -CoL--
73Shadowthrive worm Allolobophora umbrophilaAllolobophora umbrophilaKhumsan, Karjantau mountains, Tashkent Region[95][98]2 (VU:R)

Vulnerable, naturally rare

 -CoL--
74Earthworm Allolobophora graciosaAllolobophora graciosaSidzhak, Karjantau mountains, Tashkent Region[95][99]--CoL--
75Earthworm Allolobophora stenosomaAllolobophora stenosomaChimgan, Burchmullo, Sidzhak, Tashkent Region[95][100]--CoL--

Notes

  1. ↑ One Earth uses WWF ecoregions as the underlying ecological units for its bioregional classification system.

References

  1. ↑ "Mountains of Central Asia". www.cepf.net. Retrieved 2026-04-15.
  2. ↑ "Alai-Western Tian Shan Steppe". One Earth. 2022-07-29. Retrieved 2026-04-15.
  3. ↑ "Gissaro-Alai Open Woodlands". One Earth. 2021-12-15. Retrieved 2026-04-15.
  4. ↑ "Badghyz and Karabil Semi-Desert". One Earth. 2021-12-15. Retrieved 2026-04-15.
  5. ↑ "Central Asian Northern Desert". One Earth. 2021-12-15. Retrieved 2026-04-15.
  6. ↑ "Central Asian Southern Desert". One Earth. 2021-12-15. Retrieved 2026-04-15.
  7. ↑ "Central Asian Riparian Woodlands". One Earth. 2022-07-29. Retrieved 2026-04-15.
  8. 1 2 Nasekomye Uzbekistana [Insects of Uzbekistan] (in Russian). Tashkent: FAN Publishing house of Academy of Science of Uzbekistan. 1993. p. 6. ISBN 5-648-01576-3.
  9. ↑ Read, Helen J.; Golovatch, S.I. (1994). "A review of the Central Asian Millipede fauna" (PDF). Bulletin of the British Myriapod Group. 10: 59–70.
  10. 1 2 "Animals and Plants Unique to Uzbekistan". lntreasures.com. Retrieved 2026-04-15.
  11. ↑ "Living National Heritage - Species whose survival depends on the action of a single nation ... new website". Zoo's Print. 24 (6): 25–26. 2009-06-21.
  12. ↑ "Taxonomic Sources for Living National Treasures". lntreasures.com. Retrieved 2026-04-15.
  13. 1 2 Nazarov, Roman A.; Abduraupov, Timur V.; Shepelya, Evgeniya Yu.; Gritsina, Mariya A.; Melnikov, Daniel A.; Buehler, Matthew D.; Lapin, Jack D.; Poyarkov, Nikolay A.; Grismer, Jesse L. (2023-08-04). "The Fergana Valley Is an Isolate of Biodiversity: A Discussion of the Endemic Herpetofauna and Description of Two New Species of Alsophylax (Sauria: Gekkonidae) from Eastern Uzbekistan". Animals. 13 (15): 2516. doi:10.3390/ani13152516. ISSN 2076-2615. PMC 10417568. PMID 37570324.
  14. ↑ "Fergana Valley". cac-program.org. Retrieved 2026-04-15.
  15. ↑ "Fergana Valley, Kyrgyzstan: the Core of Central Asia – Central Asia Rally". www.centralasiarally.com. Retrieved 2026-04-15.
  16. ↑ Azimov, J., ed. (2019). "Fergana Smoky Moth Zygaena ferganae Sheljuzhko, 1941". The Red Data Book of the Republic of Uzbekistan in 2 volumes. Volume II (PDF) (5th ed.). Tashkent: Uzbekistan Academy of Sciences. State Committee for Ecology and Environmental Protection of the Republic of Uzbekistan. p. 56.
  17. ↑ Efetov, Konstantin (October 2012). "On the type series of Zygaena (Agrumenia) ferganae Sheljuzhko, 1941 Lepidoptera: Zygaenidae, Zygaeninae". Entomologist's Gazette. 63: 241–245.
  18. ↑ Kreuzberg, Alexander (1998). "Распространение и биология редкого вида нимфалид (Lepidoptera, Nymphalidae) — шашечницы-акреины (Меlitаеа acraeina Staudinger, 1886)" [Distribution and biology of rare species of Nimphalidae (Lepidoptera) Melitaea acraeina Staudinger, 1886] (PDF). Bulletin of Moscow Society of Naturalists. Biological Series (in Russian). 103 (1). Moscow, Russia: Publishing House of Lomonosov Moscow State University: 38–40.
  19. ↑ Naydenov, A.E.; et al. (2025-04-26). "Materials on the Lepidoptera of Uzbekistan. Part 1. Family Geometridae Leach, 1815". Acta Biologica Sibirica. 11: 589–611.
  20. ↑ "Uzbekistan Factsheet". BirdLife DataZone. Retrieved 2026-04-15.
  21. ↑ "Fergana Even-toed Gecko Alsophylax ferganensis". IUCN Red List. Retrieved 20 April 2026.
  22. ↑ "Alsophylax ferganensis". The Reptile Database. Retrieved 2026-04-20.
  23. ↑ "Alsophylax emilia". The Reptile Database. Retrieved 2026-04-20.
  24. ↑ Nazarov, Roman A.; Abduraupov, Timur V.; Shepelya, Evgeniya Yu.; Gritsina, Mariya A.; Melnikov, Daniel A.; Buehler, Matthew D.; Lapin, Jack D.; Poyarkov, Nikolay A.; Grismer, Jesse L. (2023). "The Fergana Valley Is an Isolate of Biodiversity: A Discussion of the Endemic Herpetofauna and Description of Two New Species of Alsophylax (Sauria: Gekkonidae) from Eastern Uzbekistan". Animals. 13 (15) 2516. MDPI. doi:10.3390/ani13152516. PMC 10417568. PMID 37570324.
  25. ↑ "Teratoscincus rustamowi". IUCN Red List. Retrieved 20 April 2026.
  26. ↑ Teratoscincus rustamowi at the Reptarium.cz Reptile Database
  27. ↑ IUCN (2016-04-05). Phrynocephalus reticulatus: Nuridjanov, D., Nazarov, R., Shestopal, A. & Ananjeva, N.B.: The IUCN Red List of Threatened Species 2017: e.T47755693A128947879 (Report). International Union for Conservation of Nature. doi:10.2305/IUCN.UK.2017-2.RLTS.T47755693A47755699.en.
  28. ↑ Sheraliev, Bakhtiyor; Nazirov, Bokhodirjon; Khalimov, Sharofiddin (2025-06-04). "Notes on distribution of Triplophysa ferganaensis Sheraliev & Peng, 2021 (Teleostei: Nemacheilidae)". Ecologica Montenegrina. 86: 188–196. doi:10.37828/em.2025.86.13. ISSN 2336-9744.
  29. ↑ Sheraliev, Bakhtiyor; Kayumova, Yorkinoy; Peng, Zuogang (2022-10-19). "Triplophysa daryoae, a new nemacheilid loach species (Teleostei, Nemacheilidae) from the Syr Darya River basin, Central Asia". ZooKeys (1125): 47–67. Bibcode:2022ZooK.1125...47S. doi:10.3897/zookeys.1125.85431. ISSN 1313-2970. PMC 9836642. PMID 36761291.
  30. ↑ "Triplophysa daryoae summary page". FishBase. Retrieved 2026-04-18.
  31. ↑ Sheraliev, Bakhtiyor; Kayumova, Yorkinoy (2024-11-01). "Dzihunia pseudoamudarjensis, a new nemacheilid loach from the Syr Darya drainage (Teleostei: Nemacheilidae)". Zootaxa. 5536 (1): 185–192. doi:10.11646/zootaxa.5536.1.8. ISSN 1175-5334. PMID 39646380.
  32. ↑ "Dzihunia pseudoamudarjensis summary page". FishBase. Retrieved 2026-04-18.
  33. ↑ "Dzihunia ilan". IUCN Red List. Retrieved 20 April 2026.
  34. ↑ "Orthoptera Species File - Conophyma turkestanicum Sergeev, 1984". orthoptera.speciesfile.org. Retrieved 2026-04-19.
  35. ↑ Sergeev (1984). "To the Knowledge of the Middle-Asiatic species of the genus Conophyma Zub. (Orthoptera: Acrididae)". Entomologicheskoe Obozrenie. 63 (4): 733–736.
  36. ↑ Blyummer, A.G. (2017). "A new genus and species of Paedembiidae (Embioptera) from Kyzylkum Desert (Uzbekistan)" (PDF). Russian Entomological Journal. 26 (1): 31–34.
  37. ↑ "Embioptera Species File - Uranembia rivkusi Blyummer, 2017". embioptera.speciesfile.org. Retrieved 2026-05-20.
  38. ↑ Lazarev M.A. (2009-06-28). "New species of Longicorn-beetle (Coleoptera: Cerambycidae) of the genus Neoplocaederus Sama, 1991 from Uzbekistan" (PDF). Caucasian Entomological Bulletin. 5 (1): 73–76. doi:10.23885/1814-3326-2009-5-1-73-74.
  39. ↑ "Reitterelater kovalenkoi". www.gbif.org. Retrieved 2026-04-19.
  40. ↑ Dr. Azimov J., ed. (2019). "Fergana Lethrus Lethrus bispinus (B. Jakovlev, 1899)". The Red Data Book of the Republic of Uzbekistan. Volume II (5th ed.). Tashkent: Uzbekistan Academy of Sciences. State Committee for Ecology and Environmental Protection of the Republic of Uzbekistan. p. 56.
  41. ↑ Bagaturov, M.F.; Nikolajev, G.V. (2015-11-28). "Overview of distribution of the genus Lethrus Scopoli, 1777 (Coleoptera: Geotrupidae)". Caucasian Entomological Bulletin. 11 (2): 303–314. doi:10.23885/1814-3326-2015-11-2-303-314.
  42. ↑ Bekchanov, Norbek; Nabozhenko, Maxim V.; Bekchanova, MokHira K. (2023-11-24). "Review of the subgenus Turcmenicola Bogatchev, 1952, the genus Colposcelis Dejean, 1834 (Coleoptera: Tenebrionidae: Tentyriini)". Zootaxa. 5375 (4): 550–564. doi:10.11646/zootaxa.5375.4.6. PMID 38220804.
  43. ↑ Bekchanov, Norbek; Nabozhenko, Maxim V.; Bekchanova, MokHira K. (2023-11-24). "Review of the subgenus Turcmenicola Bogatchev, 1952, the genus Colposcelis Dejean, 1834 (Coleoptera: Tenebrionidae: Tentyriini)". Zootaxa. 5375 (4): 550–564. doi:10.11646/zootaxa.5375.4.6. PMID 38220804.
  44. 1 2 3 4 5 Volkovich, M. G. (2013-08-16). "Annotated Catalogue of Jewel Beetles (Buprestidae) of the Fauna of the Former USSR". Beetles (Coleoptera) and Coleopterologists Project by Zoological Institute of the Russian Academy of Sciences, Saint Petersburg — zin.ru (in Russian).
  45. ↑ Shavrin, Alexey (September 2021). "A new species and new records of Lesteva Latreille, 1797 from the Chatkal Mountain Range, Uzbekistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)". Baltic Journal of Coleopterology. 21 (1): 49–52.
  46. ↑ Caldara R.; Korotyaev Boris A. (2002). "Description of two new species of the weevil genus Gymnetron Schoenherr, 1825 from eastern Caucasus and Uzbekistan (Coleoptera, Curculionidae)". Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano. 143: 142–145.
  47. ↑ Dr. Azimov J., ed. (2019). "Acraea Checker Spot Melitaea acraeina (Staudinger, 1886)". The Red Data Book of the Republic of Uzbekistan. Volume II (5th ed.). Tashkent: Uzbekistan Academy of Sciences. State Committee for Ecology and Environmental Protection of the Republic of Uzbekistan. p. 56.
  48. ↑ Dr. Azimov J., ed. (2019). "Fergana Smoky Moth Zygaena ferganae (Sheljuzhko, 1941)". The Red Data Book of the Republic of Uzbekistan. Volume II (5th ed.). Tashkent: Uzbekistan Academy of Sciences. State Committee for Ecology and Environmental Protection of the Republic of Uzbekistan. p. 56.
  49. ↑ "Cossulus nikiforoviorum Yakovlev, 2006". www.gbif.org. Retrieved 2026-04-19.
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  97. ↑ "Allolobophora chlorocephala Perel, 1977". WoRMS - World Register of Marine Species - www.marinespecies.org. Retrieved 2026-05-17.
  98. ↑ "Allolobophora umbrophila Perel, 1977". WoRMS - World Register of Marine Species - www.marinespecies.org. Retrieved 2026-05-17.
  99. ↑ "Allolobophora graciosa Perel, 1977". WoRMS - World Register of Marine Species - www.marinespecies.org. Retrieved 2026-05-17.
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