Cerin Lagerstätte

Cerin Lagerstätte
Stratigraphic range: Late Kimmeridgian
~
Cast of Crocodilaemus robustus from the paleontological site of Serin, showcased in the Museum of Natural Sciences of Belgium.
TypeFormation
Lithology
PrimaryLithographic limestone
Location
Coordinates45°46′44″N 05°33′15″E / 45.77889°N 5.55417°E / 45.77889; 5.55417
RegionAuvergne-Rhône-Alpes
CountryFrance
Type section
Named forCerin
Year defined1838
Cerin Lagerstätte is located in France
Cerin Lagerstätte
Cerin Lagerstätte (France)

The Cerin Lagerstätte is a fossil deposit of the Jura Mountains located in Cerin, a hamlet belonging to the commune of Marchamp in the department of Ain. The site is internationally known for its surprising diversity.[1]

The deposits are located in what was a tropical lagoon at the end of the Kimmeridgian age (Late Jurassic). It is dated from the Aulacostephanus pseudomutabilis biozone, whose equivalent in the Mediterranean area is the Aulacostephanus eudoxus biozone, which dates to 153 Ma.[2][3]

Situation

The site belonging to the Bugey historical region was erected at 560 meters above the sea level and is located 20 km off Belley, 75 km off Lyon, 80 km off Grenoble and 90 km off Geneva.[4]

Lithographic limestone

Artist view:
A Jurassic European lagoon.

Cerin was reputed by the end of the 19th century for the quality of its lithographic limestone. The area was, during the Late Jurassic, a tropical lagoon. Lithographic limestone is formed by sedimentation of a very thin carbonated mud deposited at the bottom of a lagoon 153 million of years ago. Those deposits are disposed in strata.

The quarry exploitation, which began in 1835 during the lithography golden age, allowed to periodically uncover prints of prehistoric animals and plants fossilized in stone. Paleontology was at its beginning and those discoveries were still little-known.[3]

Site discovery

A lithographic plate from « Descriptions des poissons fossiles provenant des gisements coralliens du Jura dans le Bugey », by Victor Thiollière.

In 1838, thanks to the engineer Aimé Drian, a passionate amateur geologist, and Lyonese geologist, and Victor Thiollière, those fossils were discovered and the existence of the paleontological site of Cerin was revealed to the scientific world. The site has an international reputation and rivals with the Solnhofen Limestone, in Bavaria. Until his death, Victor Thiollière never ceased to collect and study a maximum of fossils from the site, and it was, in large part, thanks to his study of the Cerin fossils that he was recognized in the palaeontology world.

His works showed the similarities of the lithographic limestones of Cerin and Solnhofen. He described several new species of fish. He published in 1854 the first part of his "Description des poissons fossiles provenant des gisements coralliens du Jura dans le Bugey" ("Description of the fossil fish from the Jura corallian deposits in the Bugey"), but died shortly before the publication of the second part, the descriptions and lithographic plates already finished.[5]

Research on site

Turtle tracks on a Japanese beach.

Research on the Late Kimmeridgian site (Late Jurassic, around −153 Ma),[6] occurred from 1975 to 1995, it was a unique operation and involved advanced technology. Directed by geologists from Claude Bernard University in Lyon, it necessitated heavy equipment of civil engineering.

This operation allowed the discovery of algae, ferns, conifers, molluscs, sea urchins, starfish, crustaceans such as the holotypes of Cyclerion bourseaui and Soleryon amicalis,[7] reptiles, fish, as well as tracks of turtles and other reptiles.[8] A thorough study permitted to determine the age and nature of the site (a tropical lagoon 153 million years old) and to understand the reasons of this exceptional fossilization.[1]

A one-of-a-kind Late Jurassic fossilized track of prehistoric marine turtle was discovered in Cerin.

Fossilization process in Cerin

Aerial view of the Aldabra atoll and its lagoon.

To understand the fossilization process in Cerin, scientists went on an expedition to Aldabra in the Indian Ocean, to observe a fossilization process similar to the one that happened on the Cerin site.

153 million years ago, the climate was tropical. The lagoon had only few contacts with the open sea, and evaporation was intense. Terrestrial animals venturing in the lagoon shores left their tracks in a quickly drying mud.

During storms, a large quantity of saltwater, carrying mud, vegetal remains, and dead or alive animals, entered the lagoon; clearwater, brought by rains and water runoff, brought as well large quantities of particulate matters. When the calm was settling, the fine particles deposited in a regular layer which carpeted the depths and covered remains and tracks.

This layer gave then birth to a strata of lithographic limestone.

When evaporation made the water level drop again, it became under oxygenated and over-concentrated on salt, which led to the death of many living beings, while protecting their bodies from scavengers. The microbial mats, who thrived in this environment, covered the corpses and vegetal remains, helping their conservation as fossils.[1]

Biota

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
 Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Algae[9]

Genus Species Classification Note Picture
CampbelliellaC. striataDasycladales
ClypeinaC. jurassicaDasycladalesClusters of this species attain more than 10 cm.
GoniolinaG. janetiDasycladales

Plants[9][10]

Genus Species Classification Note Picture
ApoldiaA. latifoliaCycadales
Brachyphyllum
B. desnoyersiiPinales
B. elegansPinales
B. speciosaPinales
Crossozamia[11]CycadopsidaFormerly described as Cycalacis.
CupressinoxylonC. spConiferales
CycadopterisC. jurensisPteridospermatophyta
PachypterisPteridospermatophyta
PagiophyllumP. cirinicumAraucariaceae
ZamitesZ. feneonisBennettitalesVery abundant in the site.
Z. pumilioBennettitales

Cnidarians[12]

Genus Species Classification Note Picture
BipedaliaB. cerinensisCubozoa
ParacarybdeaP. lithographicaCubozoa
ParaureliaP. cerinensisScyphozoa
P. spScyphozoa

Lophotrochozoans[13][9]

Genus Species Classification Note Picture
AspidocerasA. cf. catalaunicumAspidoceratidae
HibolitesBelemnoidea
Laevaptychus[14] L. latus A kind of Aptychus. Formerly referred as Aptychus latus.
LithacocerasL. cf. ulmenseAtaxioceratidae
L. sp. gr. siliceusAtaxioceratidae
L. spAtaxioceratidaePotentially a new species.
Nanogyra[15] N. virgula Bivalvia
Nerinea[15] N. suprajurensis Gastropoda
RhynchonellidaRhynchonellida
SubplanitesS. cf. rueppellianusPerisphinctidae
TerebratulidaeTerebratulidae

Arthropods[9][16]

Genus Species Classification Note Picture
Antrimpos A. speciosus Penaeidae
CycleryonC. bourseaui[7]Eryonidae
C. propinquus Eryonidae
Dusa D. monocera Penaeidae Its presence at Cerin might be a mistake caused by confusion due to misattribution of specimens hailing from the Solnhofen Limestone.
Eryma E. modestiformis Erymidae
Eryon E. arctiformis Eryonidae Its presence at Cerin might be a mistake caused by confusion due to misattribution of specimens hailing from the Solnhofen Limestone.
GigacerinaG. saemanniGlypheoideaOne of the largest known glypheids. Formerly referred to as Glyphea saemanni.
Glyphea G. pseudoscyllarus Glypheoidea Macrourites cerensis is a synonym.
Mecochirus M. brevimanus Glypheoidea Its presence at Cerin might be a mistake caused by confusion due to misattribution of specimens hailing from the Solnhofen Limestone. This species is also considered a synonym of M. longimanatus, likely to represent a female morphotype.[17]
M. spGlypheoidea
MesolimulusM. walchiLimulidae
Soleryon [7]S. amicalisEryonidae

Echinoderms[9][18][19]

Genus Species Classification Note Picture
Acrocidaris[15] A. nobilis Euechinoidea
?Ceramaster? C. chantreiGoniasteridae
Cidaris[15] C. carinifera Cidaroida
ComptoniasterC. meyeriGoniasteridae
OphiopetraO. lithographicaAplocomidae
PentasteriaP. lithographicaAstropectinidae
PseudodiademaP. cf. pseudodiademaPseudodiadematidae
PygurusClypeidae
SolanocrinitesS. thiollieriCrinoidea

Chondrichthyes[20]

Genus Species Classification Note Picture
BelemnobatisB. sismondaeAsterodermidae
CorysodonC. cirinensisCarcharhiniformesThe genus Corysodon has been synonymized with Palaeoscyllium by Cappetta, 1987. However the validity of Corysodon was maintained by Thies & Candoni, 1998[21], this was related by later studies.
PhorcynisP. catulinaSquatiniformes
SpathobatisS. bugesiacusAsterodermidae

Sarcopterygians[20]

Genus Species Classification Note Picture
UndinaU. cirinensisLatimeriidaeThis species was classified as a member of the genus Holophagus by Wenz et al., 1994[20]. However, Ferrante & Cavin, 2025[22] considered H. gulo as the only Holophagus species.

Actinopterygians[20]

Genus Species Classification Note Picture
Ainia A. armata[23] Halecomorphi
Allothrissops A. regleyi Ichthyodectiformes Formerly placed in the genus Thrissops.
Amblysemius[24] A. bellicianus Caturidae
Anaethalion A. affinis Elopiformes
A. cirinensis Elopiformes
A. knorri Elopiformes
A. cf. subovatus Elopiformes
Ankylophorus A. similis Ankylophoridae Formerly placed in the genus Pholidophorus.
Apomesodon[25] A. comosus Pycnodontiformes Synonym of A. gibbosus.
A. gibbosusPycnodontiformesFormerly referred to as Eomesodon gibbosus.[26]
A. surgens Pycnodontiformes Synonym of A. gibbosus.
Aspidorhynchus[27] A. acutirostris Aspidorhynchidae
Belonostomus B. tenuirostris Aspidorhynchidae Some authors are more prudent about the identification of this species as B. tenuirostris.[28]
B. sp.[28] Aspidorhynchidae At least one more species is present in Cerin, it is similar to B. kochii.
Caturus C. furcatus Caturidae
Cerinichthys[29] C. koelblae Halecomorphi
Eoprotelops E. vireti Elopiformes
Gyrodus "G". hexagonus[30] Pycnodontiformes The specimens from Cerin probably belong to a more derived pycnodontiform, it might not be the same species as Gyrodus hexagonus.
G. sp. Pycnodontiformes Specimens from Cerin were formerly referred to as Mesturus verrucosus.[25]
Ionoscopus I. desori Amiiformes
Lehmanophorus[31] L. segusianus Ankylophoridae Formerly referred to as Pholidophorus segusianus.
Leptolepides[32] L. sp. Orthogonikleithridae There might be more than one species present in Cerin.
Macromesodon[33] M. macropterus Pycnodontiformes Formerly referred to as Coelodus jourdani, which is now considered a junior synonym.
MacrosemiusM. fournetiMacrosemiidae
Notagogus N. helenae Macrosemiidae
N. inimontis Macrosemiidae
Oligopleurus O. esocinus[29] Halecomorphi
Ophiopsiella[34] O. guigardi Halecomorphi Formerly referred to as Ophiopsis guigardi.
O. macrodus Halecomorphi Formerly referred to as Ophiopsis macrodus.
O. procera Halecomorphi Specimens from Cerin were formerly referred to as Ophiopsis attenuata.
Ophiopsis[34]O. muensteriIonoscopiformesThe occurrence from Cerin was formerly described as Furo praelongus. However, this species has been synomymized with Furo muensteri.[35] This species was moved to the genus Ophiopsis after a taxonomic reassessment.
Orthocormus O. teyleri Pachycormiformes
Orthogonikleithrus[32] O. francogalliensis Orthogonikleithridae Specimens previously assigned to Leptolepides sprattiformis belong to this species.
O. sp. Orthogonikleithridae There is at least one more species of Orthogonikleithrus. Specimens previously referred to Ascalabos voithi actually belong to O. sp.[36]
Palaeomacrosemius[37] P. thiollieri Macrosemiidae
Pholidophorichthys P. inermis Pholidophoridae
Pholidophoristion P. cf. ovatus Pholidophoridae
"Pholidophorus"[24] "P". velifer Caturidae Formerly known as Caturus velifer, it surely does not belong to Caturus. Its taxonomic statue is still unclear and is being studied.
Pleuropholis P. thiolleri Pleuropholidae
Propterus P. microstomus Semionotiformes Material from Cerin was previously described as Histionotus falsani. This species was later regarded as a synonym of Histionotus oberndorferi, itself synonymized with Propterus microstomus.[38]
Proscinetes[39]P. bernardiPycnodontiformes
P. distantidens Pycnodontiformes
P. elegans Pycnodontiformes
P. intermedius Pycnodontiformes
P. itieri Pycnodontiformes
P. sauvanausi Pycnodontiformes
P. thiolleri Pycnodontiformes
Scheenstia[40]S. laevisLepisosteiformesFormerly referred to as Lepidotes laevis.
Solnhofenamia[29] S. elongata Amiidae Formerly placed in the genus Urocles as U. elongatus.[41]
Tharsis T. dubius Ascalaboidiformes
Thiollierepycnodus[42]T. wagneriPycnodontiformesFormerly referred to as Gyrodus wagneri.
Thrissops[43] T. cirinensis Ichthyodectiformes
T. formosus Ichthyodectiformes
Turboscinetes[44]T. egertoniPycnodontiformesFormerly referred to as Proscinetes egertoni.

Reptiles[45][46]

Genus Species Classification Note Picture
Achelonia A. formosa Eurysternidae
Alligatorellus A. beaumonti Atoposauridae
Alligatorium A. meyeri Atoposauridae
Atoposaurus A. jourdani Atoposauridae
Crocodilaemus C. robustus Pholidosauridae Sometimes orthographied Crocodileimus.
Euposaurus[47] E. cirinensis Rhynchocephalia Sometimes orthographied as E. cerinensis. This species certainly does not belong to Euposaurus.
E. lorteti Rhynchocephalia This species certainly does not belong to Euposaurus.
E. thiollierei Lepidosauromorpha Sometimes orthographied as E. thiollieri. This species is known from a single young individual and might be a lizard.
Gavialinum G. rhodani Teleosauroidea This species is now regarded as a nomen dubium.
Homoeosaurus H. jourdani Rhynchocephalia Synonym of H. maximiliani.
H. maximiliani Rhynchocephalia
H. rhodani Rhynchocephalia
Hydropelta H. meyeri Eurysternidae
Idiochelys I. fitzingeri Eurysternidae
Kallimodon K. cerinensis Rhynchocephalia
Leptosaurus L. pulchellus Rhynchocephalia
Piocormus P. cf. laticeps Rhynchocephalia Some authors have argued that Piocormus might be the same as Sapheosaurus.[47]
Pleurosaurus P. goldfussi Pleurosauridae A rhynchocephalian particularly well adapted to an aquatic lifestyle.
Pterodactylus P. cirinensis Pterosauria Now considered an indeterminate pterosaur.
Sapheosaurus S. thiollierei Rhynchocephalia
Stelliosaurus This genus is old (Gervais, 1871) and poorly documented, for those reasons, it is considered a nomen nudum and might instead be considered a synonym of Homoeosaurus.

Museum

See also

References

  1. 1 2 3 Voix de l'Ain, ed. (2009). "Cerin Marchamp: un site de fouilles mondialement connu". voixdelain.fr. Archived from the original on 9 June 2017. Retrieved 22 February 2022.
  2. ↑ Schweigert, G. 2007b. Ammonite biostratigraphy as a toolfor dating Upper Jurassic lithographic limestones from South Germany – first results and open questions. NeuesJahrbuch für Geologie und Paläontologie Abhandlungen,245(1), 117–125
  3. 1 2 "Collection des fossiles de Cerin".
  4. ↑ IGN maps on Géoportail.
  5. ↑ "Actes du colloque « Histoire des collections», Lyon, Avril 2007: Collectionneurs et collections au XIX: Eugène Dumortier et Victor Thiollière" (PDF). www.museedesconfluences.fr..
  6. ↑ Pierre Thomas. Olivier Dequincey (ed.). "La carrière de Cerin (commune de Marchamp, Ain) et ses faciès sédimentaires". Laboratoire de Sciences de la Terre / ENS Lyon. Images de la semaine. Retrieved 30 September 2012..
  7. 1 2 3 Audo, Denis; Charbonnier, Sylvain; Schweigert, Günter; Martin, Jean-Paul Saint (19 May 2014). "New eryonid crustaceans from the Late Jurassic Lagerstätten of Cerin (France), Canjuers (France), Wattendorf (Germany) and Zandt (Germany)". Journal of Systematic Palaeontology. 12 (4): 459–479. Bibcode:2014JSPal..12..459A. doi:10.1080/14772019.2013.777809. ISSN 1477-2019.
  8. ↑ "Chantiers textes". Archived from the original on 11 July 2014. Retrieved 22 February 2022.
  9. 1 2 3 4 5 Bernier, Paul; Barale, Georges; Bourseau, Jean-Paul; Buffetaut, Eric; Gaillard, Christian; Gall, Jean-Claude; Wenz, Sylvie (2014). "The lithographic limestones of Cerin (southern Jura Mountains, France). A synthetic approach and environmental interpretation". Comptes Rendus Palevol (in French). 13 (5): 383–402. Bibcode:2014CRPal..13..383B. doi:10.1016/j.crpv.2014.01.006.
  10. ↑ Barale, Georges (1978). La paléoflore jurassique du Jura français: étude systématique, aspects stratigraphiques et paléoécologiques (in French). Alexander Doweld.
  11. ↑ Zhifeng, Gao; Thomas, Barry A. (25 September 1989). "A review of fossil cycad megasporophylls, with new evidence of Crossozamia pomel and its associated leaves from the lower permian of Taiyuan, China". Review of Palaeobotany and Palynology. 60 (3): 205–223. Bibcode:1989RPaPa..60..205Z. doi:10.1016/0034-6667(89)90044-4. ISSN 0034-6667.
  12. ↑ Gaillard, Christian; Goy, Jacqueline; Bernier, Paul; Bourseau, Jean Paul; Gall, Jean Claude; Barale, Georges; Buffetaut, Eric; Wenz, Sylvie (2006). "New Jellyfish Taxa from the Upper Jurassic Lithographic Limestones of Cerin (France): Taphonomy and Ecology". Palaeontology. 49 (6): 1287–1302. Bibcode:2006Palgy..49.1287G. doi:10.1111/j.1475-4983.2006.00592.x. ISSN 0031-0239.
  13. ↑ Énay, Raymond (June 2018). "Lithostratigraphie, biostratigraphie et faunes d'ammonites du complexe corallien d'âge kimméridgien supérieur du Jura méridional et de l'Ile Crémieu (France)" (PDF). Revue de Paléobiologie, Genève. 37 (1): 41–108.
  14. ↑ Zell, Patrick; Stinnesbeck, Wolfgang; Beckmann, Seija; Zell, Patrick; Stinnesbeck, Wolfgang; Beckmann, Seija (2016). "Late Jurassic aptychi from the La Caja Formation of northeastern Mexico". Boletín de la Sociedad Geológica Mexicana. 68 (3): 515–536. Bibcode:2016BoSGM..68..515Z. doi:10.18268/BSGM2016v68n3a8. ISSN 1405-3322.
  15. 1 2 3 4 de Saint-Seine, Pierre (1949). "Les poissons des calcaires lithographiques de Cerin (Ain)". Publications du musée des Confluences. 2: 3–79.
  16. ↑ Van Straelen, V. (1922). "LES CRUSTACES DECAPODES DU PORTLANDIEN DE CERIN-MARCHAMPT (AIN)". Comptes rendus hebdomadaires des séances de l'Académie des sciences.
  17. ↑ Schweitzer, Carrie; Feldmann, Rodney; Garassino, Alessandro; Karasawa, Hiroaki; Schweigert, Günter (2010). Systematic List of Fossil Decapod Crustacean Species. doi:10.1163/ej.9789004178915.i-222. ISBN 978-90-474-4126-7. Retrieved 1 April 2026. {{cite book}}: |website= ignored (help)
  18. ↑ Breton, Gérard; Bourseau, Jean-Paul; Bernier, Paul; Barale, Georges; Buffetaut, Éric; Gaillard, Christian; Gall, Jean-Claude; Wenz, Sylvie (1 June 1994). "Les Astérides (Asteroidea, Echinodermata) des calcaires lithographiques kimméridgiens de Cerin (Ain, France)". Geobios. 27: 49–60. Bibcode:1994Geobi..27...49B. doi:10.1016/S0016-6995(94)80020-0. ISSN 0016-6995.
  19. ↑ Bourseau, Jean-Paul; Bernier, Paul; Barale, Georges; Buffetaut, Eric; Gaillard, Christian; Gall, Jean-Claude; Roman, Jean; Wenz, Sylvie (1 June 1994). "Taphonomie des Échinides dugisement de Cerin (kimméridgien Supérieur, Jura Méridional, France). Implications environnementales". Geobios. 27: 37–47. Bibcode:1994Geobi..27...37B. doi:10.1016/S0016-6995(94)80019-7. ISSN 0016-6995.
  20. 1 2 3 4 Wenz, Sylvie; Bernier, Paul; Barale, Georges; Bourseau, Jean-Paul; Buffetaut, Eric; Gaillard, Christian; Gall, Jean-Claude (1 June 1994). "L'ichthyofaune des calcaires lithographiques du Kimméridgien supérieur de Cerin (Ain, France)". Geobios. 27: 61–70. Bibcode:1994Geobi..27...61W. doi:10.1016/S0016-6995(94)80021-9. ISSN 0016-6995.
  21. ↑ Thies, Detlev; Candoni, Laurent (1998). "Corysodon SAINT-SEINE 1949 - a valid genus of Mesozoic neoselachian sharks". Geologica et Palaeontologica.
  22. ↑ Ferrante, Christophe; Cavin, Lionel (6 June 2025). "A deep dive into the coelacanth phylogeny". PLOS ONE. 20 (6) e0320214. Bibcode:2025PLoSO..2020214F. doi:10.1371/journal.pone.0320214. ISSN 1932-6203. PMC 12143573. PMID 40478838.
  23. ↑ Rauhut, O. W. M.; López-Arbarello, A.; Röper, M.; Rothgaenger, M. (2017). "Vertebrate fossils from the Kimmeridgian of Brunn: the oldest fauna from the Solnhofen Archipelago (Late Jurassic, Bavaria, Germany)" (PDF). Zitteliana. 89: 305–329.
  24. 1 2 López-Arbarello, Adriana; Ebert, Martin (11 January 2023). "Taxonomic status of the caturid genera (Halecomorphi, Caturidae) and their Late Jurassic species". Royal Society Open Science. 10 (1) 221318. Bibcode:2023RSOS...1021318L. doi:10.1098/rsos.221318. ISSN 2054-5703. PMC 9832298. PMID 36686548.
  25. 1 2 "A new insight into pycnodontiform fishes". Geodiversitas. 24 (1): 139–248. 2002.
  26. ↑ Ebert, Martin (3 February 2026). "Macromesodon Blake, 1905 and Apomesodon Poyato-Ariza & Wenz, 2002 (Actinopterygii, Pycnodontiformes) from the Jurassic and lowermost Cretaceous of England, France, and Germany". Swiss Journal of Palaeontology. 145: 97–129. doi:10.3897/sjp.145.177263. ISSN 1664-2384.
  27. ↑ Brito, Paulo M.; Meunier, François J. (1 January 2000). "The morphology and histology of the scales of Aspidorhynchidae (Actinopterygii, Halecostomi)". Geobios. 33 (1): 105–111. Bibcode:2000Geobi..33..105B. doi:10.1016/S0016-6995(00)80153-9. ISSN 0016-6995.
  28. 1 2 Ebert, Martin (2014). "The genus Belonostomus Agassiz, 1834 (Neopterygii, Aspidorhynchiformes) in the late Jurassic of the Solnhofen Archipelago, with a focus on Belonostomus kochii Münster, 1836 from Ettling (Germany)". Archaeopteryx. 32: 15–43.
  29. 1 2 3 Ebert, Martin (2 January 2018). "Cerinichthys koelblae , gen. et sp. nov., from the Upper Jurassic of Cerin, France, and its phylogenetic setting, leading to a reassessment of the phylogenetic relationships of Halecomorphi (Actinopterygii)". Journal of Vertebrate Paleontology. 38 (1) e1420071. Bibcode:2018JVPal..38E0071E. doi:10.1080/02724634.2017.1420071. ISSN 0272-4634.
  30. ↑ Kriwet, Jürgen (25 September 2000). "Revision of Mesturus cordillera Martill et al., 1998 (Actinopterygii, Pycnodontiformes) from the Oxfordian (Upper Jurassic) of northern Chile". Journal of Vertebrate Paleontology. 20 (3): 450–455. doi:10.1671/0272-4634(2000)020[0450:ROMCME]2.0.CO;2. ISSN 0272-4634.
  31. ↑ Gaudant, Jean (1978). "Essai de révision taxonomique des « Pholidophorus » (Poissons Actinoptérygiens) du Jurassique supérieur de Cerin (Ain)". Publications du musée des Confluences. 16: 101–121.
  32. 1 2 Konwert, Martin (3 May 2016). "Orthogonikleithrus francogalliensis , sp. nov. (Teleostei, Orthogonikleithridae) from the Late Jurassic Plattenkalks of Cerin (France)". Journal of Vertebrate Paleontology. 36 (3) e1101377. Bibcode:2016JVPal..36E1377K. doi:10.1080/02724634.2016.1101377. ISSN 0272-4634.
  33. ↑ Ebert, Martin (3 February 2026). "Macromesodon Blake, 1905 and Apomesodon Poyato-Ariza & Wenz, 2002 (Actinopterygii, Pycnodontiformes) from the Jurassic and lowermost Cretaceous of England, France, and Germany". Swiss Journal of Palaeontology. 145: 97–129. doi:10.3897/sjp.145.177263.
  34. 1 2 Lane, Jennifer A.; Ebert, Martin (2 January 2015). "A taxonomic reassessment of Ophiopsis (Halecomorphi, Ionoscopiformes), with a revision of Upper Jurassic species from the Solnhofen Archipelago, and a new genus of Ophiopsidae". Journal of Vertebrate Paleontology. 35 (1) e883238. Bibcode:2015JVPal..35E3238L. doi:10.1080/02724634.2014.883238. ISSN 0272-4634.
  35. ↑ Lane, Jennifer A.; Ebert, Martin (June 2012). "Revision of Furo muensteri (Halecomorphi, Ophiopsidae) from the Upper Jurassic of Western Europe, with comments on the genus". Journal of Vertebrate Paleontology. 32 (4): 799–819. Bibcode:2012JVPal..32..799L. doi:10.1080/02724634.2012.680325. ISSN 0272-4634.
  36. ↑ Konwert, Martin (2017). "Contributions to the Taxonomy, Diversity and Palaebiogeography of certain basal Teleostei (Osteichthyes, Actinopterygii) from the Early and Late Jurassic of Central Europe". Via ResearchGate.
  37. ↑ Ebert, Martin; Lane, Jennifer A.; Kölbl-Ebert, Martina (2 September 2016). "Palaeomacrosemius thiollieri, gen. et sp. nov., a new Macrosemiidae (Neopterygii) from the Upper Jurassic of the Solnhofen Archipelago (Germany) and Cerin (France), with a revision of the genus Macrosemius". Journal of Vertebrate Paleontology. 36 (5) e1196081. Bibcode:2016JVPal..36E6081E. doi:10.1080/02724634.2016.1196081. ISSN 0272-4634.
  38. ↑ Ebert, Martin (December 2012). "Histionotus (Actinopterygii, Macrosemiidae) – Eine Gattung mit vielen Fragezeichen". Via ResearchGate.
  39. ↑ Ebert, Martin (2013). "The Pycnodontidae (Actinopterygii) in the late Jurassic: 1) he genus Proscinetes Gistel, 1848 in the Solnhofen Archipelago (Germany) and Cerin (France)". Archaeopteryx – via Research Gate.
  40. ↑ López-Arbarello, Adriana (11 July 2012). "Phylogenetic Interrelationships of Ginglymodian Fishes (Actinopterygii: Neopterygii)". PLOS ONE. 7 (7) e39370. Bibcode:2012PLoSO...739370L. doi:10.1371/journal.pone.0039370. ISSN 1932-6203. PMC 3394768. PMID 22808031.
  41. ↑ Grande, Lance; Bemis, William E. (10 April 1998). "A Comprehensive Phylogenetic Study of Amiid Fishes (Amiidae) Based on Comparative Skeletal Anatomy. an Empirical Search for Interconnected Patterns of Natural History". Journal of Vertebrate Paleontology. 18 (sup1): 1–696. Bibcode:1998JVPal..18S...1G. doi:10.1080/02724634.1998.10011114. ISSN 0272-4634.
  42. ↑ Ebert, Martin (21 October 2019). "A new genus of Pycnodontidae (Actinopterygii) from the Upper Jurassic of France and Germany, included in a phylogeny of Pycnodontiformes". Zoological Journal of the Linnean Society zlz087. doi:10.1093/zoolinnean/zlz087. ISSN 0024-4082.
  43. ↑ Ebert, Martin (25 July 2025). "New species of the genus Thrissops (Teleostei, Ichthyodectiformes) in the Upper Jurassic of the Solnhofen-Archipelago (Germany) and Kimmeridge Clay (England)". Zitteliana. 99: 1–32. doi:10.3897/zitteliana.99.159055. ISSN 2747-8106.
  44. ↑ Ebert, Martin (March 2016). "The Pycnodontidae (Actinopterygii) in the late Jurassic: 2) Turboscinetes gen. nov. in the Solnhofen Archipelago (Germany) and Cerin (France)". Via ResearchGate.
  45. ↑ Brignon, Arnaud (2025). "Les planches lithographiées des « Reptiles fossiles du Bugey (Ain) » de Claude Jourdan (1803-1873) et leur apport à l'histoire des premières découvertes de reptiles dans le Jurassique de Cerin et des gisements voisins". Revue de Paléobiologie, Genève. 44 (12): 1–57.
  46. ↑ Fabre, Jacques (1981). Les rhynchocephales et les pterosauriens a crete parietale du Kimeridgien superieur-Berriasien d'Europe occidentale: le gisement de Canjuers (Var-France) et ses abords (in French). Paris: Editions de la Fondation Singer-Polignac. ISBN 978-2-402-41853-9.
  47. 1 2 Evans, Susan E. (1 January 1994). "A re-evaluation of the Late Jurassic (Kimmeridgian) reptile Euposaurus (Reptilia: Lepidosauria) from Cerin, France". Geobios. 27 (5): 621–631. Bibcode:1994Geobi..27..621E. doi:10.1016/S0016-6995(94)80261-0. ISSN 0016-6995.

Bibliography and further reading

  • Victor Thiollière, Descriptions des poissons fossiles provenant des gisements coralliens du Jura dans le Bugey, Paris, Editions J.-B. Baillière, 1854.
  • Victor Thiollière et Paul Gervais, Descriptions des poissons fossiles provenant des gisements coralliens du Jura dans le Bugey. 2e partie. Revue et annotée par Paul Gervais avec l'aide de Gaston de Saporta, Falsan et Dumortier, Lyon, Éditions H. Georg, 1873.
  • Louis David, Une lagune tropicale au temps des dinosaures, édition du CNRS, 1985.
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